Description

Key Characters:

Growth Form: Erect or ascending perennial herbs 4–20 dm tall, sometimes somewhat woody toward base.

Stems: Stems few to many-branched, glabrous or sparsely strigillose.

Roots:

Leaves: Leaves simple. Opposite and decussate (pairs of leaves alternating 90 degrees between nodes). Blades lanceolate to oblanceolate, 3–10 cm long, 1–1.5 cm wide. Apex acute. Base attenuate. Surfaces finely scaberulous. Margins coarsely serrate. Veins prominent. Sessile. Stipules absent.

Flowers: Flowers numerous in several slender, terminal spikes 3–12 cm long, sparsely short–pilose throughout; Bracts subulate, shorter than to as long as the calyx; short–pilose. Flowers bisexual (perfect), alternate, nearly zygomorphic. Calyx sometimes purple, 1.5–2.3 mm long. Corolla usually blue, occasionally pale blue, lavender, or purple, ca. 2–3 mm long, 5-lobed, imbricate. Stamens 4, in 2 pairs, inserted on and included in corolla tube; anthers dithecal, opening by longitudinal slits. Ovary superior, 4-lobed, 2-carpellate, 4-celled, placentation axile; ovules 1 per cell, attached laterally or near base, erect, anatropous; style 2-lobed, the lobes dissimilar, only 1 stigmatic.

Fruit: Schizocarps readily separating at maturity into 4 crustaceous nutlets; pericarp hard and dry. Seeds 1 per nutlet; oblong; triangular in cross section; endosperm essentially absent.

Ploidy: 2n = 28; 56

Habitat: Occurring in dry to wet; disturbed habitats.

Elevation Range: 10–2,280 m.

Historical Distribution

Images

Synonyms (32)

Verbena affinis M.Martens & Galeotti (1844)

Verbena bonariensis (Misapplied)

Verbena bonariensis var. litoralis (Kunth) Hook. ex Müll.Berol. (1845)

Verbena brasiliensis var. subglabrata Moldenke (1950)

Verbena cajamarcensis Binder ex Molinari (2015)

Verbena caracasana Kunth (1818)

Verbena carolina var. glabra Moldenke (1959)

Verbena gentryi Moldenke (1941)

Verbena glabrata var. tenuispicata Moldenke (1967)

Verbena integrifolia Sessé & Moc. (1886)

Verbena integrifolia f. albiflora Moldenke (1968)

Verbena lanceolata Willd. ex Spreng. (1825)

Verbena litoralis f. albiflora (Moldenke) Moldenke (1979)

Verbena litoralis f. magnifolia Moldenke (1977)

Verbena litoralis var. albiflora Moldenke (1940)

Verbena litoralis var. caracasana (Kunth) Briq. (1904)

Verbena litoralis var. glabrior Benth. (1846)

Verbena litoralis var. leptostachya Schauer (1847)

Verbena litoralis var. litoralis

Verbena litoralis var. melanopotamica Hauman (1913)

Verbena litoralis var. portoricensis Moldenke (1982)

Verbena litoralis var. pycnostachya Schauer (1847)

Verbena litoralis var. subglabrata (Moldenke) N.O'Leary & Múlgura (2012)

Verbena longifolia M.Martens & Galeotti (1844)

Verbena longifolia f. albiflora Moldenke (1961)

Verbena longifolia var. pubescens Moldenke (1966)

Verbena minutiflora var. peruviana Moldenke (1981)

Verbena nudiflora Nutt. ex Turcz. (1863)

Verbena paucifolia Turcz. (1863)

Verbena sedula Moldenke (1955)

Verbena sedula var. darwinii Moldenke (1968)

Verbena sedula var. fournieri Moldenke (1969)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Ni'ihau(Incl. Lehua) Naturalized

Kuaihelani (Midway Atoll) Naturalized

Kaua'i Naturalized

O'ahu Naturalized

Molokai Naturalized

Lana'i Naturalized

Kaho'olawe Naturalized

Maui Naturalized

Hawai'i Naturalized

Dispersal Agents

Pollinators

Specimens

Conservation Status

IUCN:Data Deficient

Hawaii Pacific Weed Risk Assessment Rating:High Risk

HPWRA Score:15

Notes

Description digitized by Ikaika Mendez
Description digitized from the Manual of the Flowering Plants of Hawaii
Has been used medicinally in Hawai'i especially as a mash applied to cuts and bruises and also to sprained and fractured areas.
in 1837, specific locality unknown

Bibliography

Name Published In: F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 2: 276 (1818)

Other References

Wagner et al. 1990:1325 (Mi, Ni, K, O, Mo, L, M, Ka, H)

Occurrences

SNo.	Scientific Name	Locality	Habitat	Basis of Record	Description	Recorded By	Record Number	Island	Source	Date
1	Verbena litoralis	Lower half of Kaala Road	mesic forest	Preserved_Specimen	flowers lavender	Annable, C.R.	3727	Oahu	BISH	1997-10-15
2	Verbena litoralis	E central part of island, slope of Moaula	Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping.	Preserved_Specimen	Flowers purple.	Cuddihy, L.W.	428	Kahoolawe	BISH	1980-04-25
3	Verbena litoralis	Waikane-Schofield trail, Koolau Mts, Kahana	along trail	Preserved_Specimen	fls. lavendar	Fosberg, F.R.	10276	Oahu	BISH	1933-09-22
4	Verbena litoralis	Trail to Kaluiwaa (Sacred Falls), outside forest Reserve	At base of foot path trail.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-03-21
5	Verbena litoralis	East Maui, Haleakala Crater, Kaupo Gap	Dry volcanic gravel by trail.	Preserved_Specimen	Flws lavender-blue.	St.John, H.	21182	Maui	BISH	1945-08-27
6	Verbena litoralis	Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road	Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside..	Preserved_Specimen		Newell, C.L.	471	Hawaii	BISH	1966-08-20
7	Verbena litoralis	Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979	Pohakuloa burn 1979.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
8	Verbena litoralis	ahupuaa of Halepuaa Forest Reserve in the Puna District	Found growing on the side of the old railroad.	Preserved_Specimen		ESP Field Crew		Hawaii	BISH	1979-08-15
9	Verbena litoralis	North Kohala District, Kalala ahupua'a, Lahikiola cinder cone	In Kikuyu grassland	Preserved_Specimen		Bunch, P.M.	112	Hawaii	BISH	1989-03-13
10	Verbena litoralis	Kipapa Gulch, Wahiawa	moderately moist pineapple field	Preserved_Specimen		Hosaka, E.Y.	422	Oahu	BISH	1931-06-18
11	Verbena litoralis	Waimea Audubon Center, Waimea Valley, Oahu, Hawaii	Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions.	Preserved_Specimen	Shrub, spindly, flowers purple, on spikes.	Crago, L.M.	2005207	Oahu	BISH	2005-10-18
12	Verbena litoralis	Waiapaa Gulch, flat area near the road above	Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants.	Preserved_Specimen		Funk, E.J.	920004	Lanai	BISH	1992-01-26
13	Verbena litoralis	South Kohala: Waimea, Lalamilo	Range paddock; occasional.	Preserved_Specimen	Branching plant 1-3 ft.	Hosaka, E.Y.	2122	Hawaii	BISH	1938-06-01
14	Verbena litoralis	Kaalualu Bay near South Point	Only one individual found in waste area.	Preserved_Specimen	Herb up to one m in height with small purplish flowers.	Whistler, W.A.	42	Hawaii	BISH	1972-04-06
15	Verbena litoralis	Lawai Valley; P.T.B.G		Preserved_Specimen	Herb w/pale blue flowers. Uncommon in valley.	Herbst, D.R.	2296	Kauai	BISH	1972-01-04
16	Verbena litoralis	East Maui, Upper Kaupo Gap	in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground.	Preserved_Specimen	Flowers purple	Higashino, P.K.	885	Maui	BISH	1975-06-22
17	Verbena litoralis	Located on the Mauna Loa Strip road	about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19	Preserved_Specimen	Shrub about 4 ft. tall	Newell, C.L.	966	Hawaii	BISH	1967-06-23
18	Verbena litoralis	near Koko Head, Kaluanui Ridge,Koolau Range	guava zone; at summit	Preserved_Specimen		Egler, F.E.	3783	Oahu	BISH	1937-07-01
19	Verbena litoralis	Kaimuki, Honolulu	roadside	Preserved_Specimen		Fosberg, F.R.	8856	Oahu	BISH	1932-09-24
20	Verbena litoralis	0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline;	on rockpile, vegetation Myoporum-Chamaesyce dominant,	Preserved_Specimen		Wagner, W.L.	4688	Hawaii	BISH	1982-11-17
21	Verbena litoralis	Growing next to Kaluanui Stream	Approximately 1/2 mile from ocean.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-05-18
22	Verbena litoralis	Waianae Mts., Makaha Valley	on trail	Preserved_Specimen		Hasegawa, M.		Oahu	BISH	1932-04-03
23	Verbena litoralis	Keauhou Ranch, Kau District		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/16/2007
24	Verbena litoralis	Kalaoa, HI 96740, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	10/18/2007
25	Verbena litoralis	Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field	Roadside.	Preserved_Specimen		Wagner, W.L.	4826	Oahu	BISH	1983-04-04
26	Verbena litoralis	Honolulu, Hawaii, United States		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
27	Verbena litoralis	Growing next to Kaluanui Stream		OBSERVATION		Curtis Daehler		Oahu	HAPI	3/21/2007
28	Verbena litoralis	S half of island	Ponds	Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
29	Verbena litoralis	East Maui, floor of Haleakala Crater	1/2 mile East of Holua Cabin along trail	Preserved_Specimen		Henrickson, J.S.	3939	Maui	BISH	1969-07-20
30	Verbena litoralis	Maui		Preserved_Specimen	Flower lavender	Handy, E.S.	3320	Maui	BISH	1933-06-10
31	Verbena litoralis	Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ)	Common roadside weed.	Preserved_Specimen		Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
32	Verbena litoralis	ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision	The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica.	Preserved_Specimen		Yoshida, L.	79.053	Hawaii	BISH	1979-02-28
33	Verbena litoralis	Wahiawa		Preserved_Specimen	Erect herb, 1m; flower purple	Swarbrick, J.T.	0.62	Oahu	BISH	1976-01-07
34	Verbena litoralis	Haena		Preserved_Specimen				Kauai	BISH	1953-10-30
35	Verbena litoralis	Weed of pineapple field near northern junction of Highways 37 and 377		Preserved_Specimen	Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple.	Herbst, D.R.	795	Maui	BISH	1967-12-18
36	Verbena litoralis	Kanehoa, Waianaeuka	near top of ridge	Preserved_Specimen		Hume, E.P.	123	Oahu	BISH	1931-03-22
37	Verbena litoralis	East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store	Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus.	Preserved_Specimen		Imada, C.	200292	Maui	BISH	2002-11-19
38	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen	Herb, woody base,1.25 m high	Christophersen, E.		Oahu	BISH	1924-12-07
39	Verbena litoralis	Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility		Preserved_Specimen	Herb 1 m tall; flws light violet.	Lammers, T.G.	5313	Oahu	BISH	1983-08-27
40	Verbena litoralis	Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail		Preserved_Specimen		Wagner, W.L.	5398	Molokai	BISH	1984-07-17
41	Verbena litoralis	Maui		Preserved_Specimen		Forbes, C.N.	2036	Maui	BISH	1920-04-19
42	Verbena litoralis	Hawaii County, HI, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
43	Verbena litoralis	Pohakuloa dry area		Preserved_Specimen		Long, C.R.	4309	Hawaii	BISH	1966-12-23
44	Verbena litoralis	Maui		Preserved_Specimen		Wagner, W.L.	4783	Maui	BISH	1983-01-14
45	Verbena litoralis	Oahu		Preserved_Specimen		Rémy, J.	406	Oahu	BISH	1851-01-01
46	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	141	Lanai	BISH	1913-10-10
47	Verbena litoralis	Area within 300 m west of LZ-1. UTM coordinates 752400-2275900		Preserved_Specimen		Warren, S.	14	Kahoolawe	BISH	1987-04-09
48	Verbena litoralis	Waipio		Preserved_Specimen		Swanson, R.		Hawaii	BISH	1931-08-29
49	Verbena litoralis	Tantalus Rd		Preserved_Specimen		Forbes, A.		Oahu	BISH	1930-07-18
50	Verbena litoralis	Waikolu Valley, head of and E. rim		Preserved_Specimen		Kondo, Y.		Molokai	BISH	1962-06-18
51	Verbena litoralis	Honolulu		Preserved_Specimen		Pukui, M.K.		Oahu	BISH	1931-01-01
52	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen		Christophersen, E.		Oahu	BISH	1924-12-07
53	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
54	Verbena litoralis	Waianae Mts., South end		Preserved_Specimen		Harley, K.		Oahu	BISH	1956-05-04
55	Verbena litoralis	Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea		Preserved_Specimen		Adee, K.T.		Hawaii	BISH
56	Verbena litoralis	Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road		Preserved_Specimen		Ziegler, M.F.		Oahu	BISH	1986-11-08
57	Verbena litoralis	Kealia Salt Marsh		Preserved_Specimen		Funk, E.J.		Maui	BISH	1990-02-15
58	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
59	Verbena litoralis	Mokuleia Forest Reserve, Waianae Mts		Preserved_Specimen		Onouye, R.		Oahu	BISH	1934-02-04
60	Verbena litoralis	Valley above country club, Honolulu		Preserved_Specimen		Forbes, C.N.		Oahu	BISH	1908-07-08
61	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
62	Verbena litoralis	Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above		Preserved_Specimen		Sohmer, S.H.	5995	Oahu	BISH	1969-08-27
63	Verbena litoralis	Lanai		Preserved_Specimen		Munro, G.C.	292	Lanai	BISH
64	Verbena litoralis	Slopes of Mauna Kea (Wailei)		Preserved_Specimen		Forbes, C.N.	465	Hawaii	BISH	1911-08-01
65	Verbena litoralis	Kaua Trail, Waianae Mts		Preserved_Specimen		Woolford, H.A.	200	Oahu	BISH	1955-12-11
66	Verbena litoralis	Kealia trail		Preserved_Specimen		Bennett, T.	244	Oahu	BISH	1982-03-25
67	Verbena litoralis	Kualoa Pasture		Preserved_Specimen		Rock, J.F.C.	3007	Oahu	BISH	1942-01-01
68	Verbena litoralis	Kaholuamanoa, above Waimea		Preserved_Specimen		Heller, A.A.	2046	Kauai	BISH	1895-09-02
69	Verbena litoralis	Pauoa Valley		Preserved_Specimen		Garber, D.W.	339	Oahu	BISH	1920-03-21
70	Verbena litoralis	Honolulu, Aihualama		Preserved_Specimen		Russell, P.	253	Oahu	BISH	1932-07-08
71	Verbena litoralis	Crater of Haleakala		Preserved_Specimen		Forbes, C.N.	1083	Maui	BISH	1919-08-09
72	Verbena litoralis	Koele		Preserved_Specimen		Forbes, C.N.	188	Lanai	BISH	1918-06-01
73	Verbena litoralis	Maui		Preserved_Specimen		Neal, M.C.		Maui	BISH	1933-08-01
74	Verbena litoralis	unknown		Preserved_Specimen		Mann, H.	437		BISH	1864-01-01
75	Verbena litoralis	Kilauea, Kaloko Reservoir		Preserved_Specimen		Forbes, C.N.	572	Kauai	BISH	1916-10-11
76	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	365	Lanai	BISH	1928-08-01
77	Verbena litoralis	Palolo Valley, middle ridge	forest	Preserved_Specimen		Christophersen, E.	1666	Oahu	BISH	1931-03-26
78	Verbena litoralis	Palaau, west side of airport	pineapple field; dry, exposed, windy.	Preserved_Specimen	Herb to sub-shrub with blue flowers	Herbst, D.R.	5092	Molokai	BISH	1974-11-11
79	Verbena litoralis	Mt. Kalena, Waianae	Lantana grove	Preserved_Specimen	herb 4 ft.	Grant, M.L.	7445	Oahu	BISH	1934-09-30
80	Verbena litoralis	Koloa, Kalaheo, upper end of Puuwai Rd.	wet roadside secondary forest; Wedelia	Preserved_Specimen	to 0.5 m herb, slender; square stem, small purple flowers	Trauernicht, C.	205	Kauai	BISH	2007-10-15
81	Verbena litoralis	Nawaimaka drainage.	Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai	Preserved_Specimen	1.5 m. tall, stems green, corolla pale pruple.	Wood, K.R.	13816	Kauai	BISH	2009-07-16
82	Verbena litoralis	ridge north of Kuapa Pond	open, hillsides	Preserved_Specimen		Gillespie, J.W.	1076	Oahu	BISH	1927-07-08
83	Verbena litoralis	Kau District: Keauhou Ranch	Along fenceline of horse corrall in lower pastures of ranch.	Preserved_Specimen	Herb, 3 ft. tall, purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
84	Verbena litoralis	stream bank below Dillingham Field wetland	grasses	Preserved_Specimen		Funk, E.J.	1105	Oahu	BISH	1990-05-08
85	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
86	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
87	Verbena litoralis	Makanikahau Reservoir, Old	Dried mud	Preserved_Specimen		St.John, H.	23633	Niihau	BISH	1949-04-01
88	Verbena litoralis	Puu Huluhulu. Near the escarpment Transect (T16)	Remnant dryland community. Common subshrub among Pennisetum setaceum.	Preserved_Specimen	Common subshrub.	Takeuchi, W.N.	5714	Hawaii	BISH	1989-12-26
89	Verbena litoralis	Kaluanui Flats	Pineapple fields	Preserved_Specimen	Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas.	Spence, G.	369	Lanai	BISH	1973-10-24
90	Verbena litoralis	Mauna Kea (Puu Laau Road)	In open Mamane-Naio forest.	Preserved_Specimen		Engilis Jr., A.	39	Hawaii	BISH	1987-08-19
91	Verbena litoralis	Keanakolu, Hilo Forest Reserve	Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus.	Preserved_Specimen		Higashino, P.K.	6965	Hawaii	BISH	1978-03-01
92	Verbena litoralis	Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost	in open sunny area.	Preserved_Specimen		Herat, T.	559	Hawaii	BISH	1974-06-12
93	Verbena litoralis	North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary	Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa	Preserved_Specimen		Close, B.	109426	Hawaii	BISH	1994-10-29
94	Verbena litoralis	Peacock Flat, Waianae Mts	roadside, dry forest	Preserved_Specimen	herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue.	Spence, G.	209	Oahu	BISH	1973-09-06
95	Verbena litoralis	Puu Laau	Weed growing near the hunter's cabin.	Preserved_Specimen		Herbst, D.R.	5210	Hawaii	BISH	1975-01-18
96	Verbena litoralis	Kahaluu; Kailua; Kapua	Koa forest.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
97	Verbena litoralis	Puuwaawaa; track to 1986 burn area (from Ranch HQ)	4WD access track to burn area.	Preserved_Specimen	Common roadside weed.	Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
98	Verbena litoralis	Kawa side of Kahana Valley, pass the main gate and the marshy area	along jeep trail, open areas	Preserved_Specimen	common weed, ~4', with light blue flowers	Herat, T.	158	Oahu	BISH	1972-08-14
99	Verbena litoralis	1750' Field 123, Honokaa GSugar Co. Hamakua District	Common weed at field edge, and on jeep roads through field, not present inside mature stand.	Preserved_Specimen	Perennial root, plants 30- 48 tall, corolla pale blue	Tomich, P.Q.	911	Hawaii	BISH	1960-03-18
100	Verbena litoralis	Kahuku, Ocean View Estates,	open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive.	Preserved_Specimen	Common roadside weed with blue flowers.	Herbst, D.R.	5068	Hawaii	BISH	1974-10-09
101	Verbena litoralis	Kilauea	weed in abandoned sorghum field	Preserved_Specimen		Herbst, D.R.	5244	Kauai	BISH	1975-02-27
102	Verbena litoralis	Laie	clay, pasture	Preserved_Specimen	herb 1-6', tall slender stems	Guinto, J.	27	Oahu	BISH	1986-03-06
103	Verbena litoralis	Honolulu, 3830 Tantalus Dr., access road to Puu Ohia	roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum.	Preserved_Specimen	to 4-20 dm tall; corolla blue	Ray, G.	50	Oahu	BISH	1997-09-15
104	Verbena litoralis	Kewalo, back of Punchbowl	open, dry, roadside	Preserved_Specimen	1 m tall; flower - small, inconspicous, lavender	Judd, C.S.	11	Oahu	BISH	1924-11-06
105	Verbena litoralis	Lower half of Kaala Road	mesic forest	Preserved_Specimen	flowers lavender	Annable, C.R.	3727	Oahu	BISH	1997-10-15
106	Verbena litoralis	E central part of island, slope of Moaula	Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping.	Preserved_Specimen	Flowers purple.	Cuddihy, L.W.	428	Kahoolawe	BISH	1980-04-25
107	Verbena litoralis	Waikane-Schofield trail, Koolau Mts, Kahana	along trail	Preserved_Specimen	fls. lavendar	Fosberg, F.R.	10276	Oahu	BISH	1933-09-22
108	Verbena litoralis	Trail to Kaluiwaa (Sacred Falls), outside forest Reserve	At base of foot path trail.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-03-21
109	Verbena litoralis	East Maui, Haleakala Crater, Kaupo Gap	Dry volcanic gravel by trail.	Preserved_Specimen	Flws lavender-blue.	St.John, H.	21182	Maui	BISH	1945-08-27
110	Verbena litoralis	Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road	Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside..	Preserved_Specimen		Newell, C.L.	471	Hawaii	BISH	1966-08-20
111	Verbena litoralis	Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979	Pohakuloa burn 1979.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
112	Verbena litoralis	ahupuaa of Halepuaa Forest Reserve in the Puna District	Found growing on the side of the old railroad.	Preserved_Specimen		ESP Field Crew		Hawaii	BISH	1979-08-15
113	Verbena litoralis	North Kohala District, Kalala ahupua'a, Lahikiola cinder cone	In Kikuyu grassland	Preserved_Specimen		Bunch, P.M.	112	Hawaii	BISH	1989-03-13
114	Verbena litoralis	Kipapa Gulch, Wahiawa	moderately moist pineapple field	Preserved_Specimen		Hosaka, E.Y.	422	Oahu	BISH	1931-06-18
115	Verbena litoralis	Waimea Audubon Center, Waimea Valley, Oahu, Hawaii	Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions.	Preserved_Specimen	Shrub, spindly, flowers purple, on spikes.	Crago, L.M.	2005207	Oahu	BISH	2005-10-18
116	Verbena litoralis	Waiapaa Gulch, flat area near the road above	Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants.	Preserved_Specimen		Funk, E.J.	920004	Lanai	BISH	1992-01-26
117	Verbena litoralis	South Kohala: Waimea, Lalamilo	Range paddock; occasional.	Preserved_Specimen	Branching plant 1-3 ft.	Hosaka, E.Y.	2122	Hawaii	BISH	1938-06-01
118	Verbena litoralis	Kaalualu Bay near South Point	Only one individual found in waste area.	Preserved_Specimen	Herb up to one m in height with small purplish flowers.	Whistler, W.A.	42	Hawaii	BISH	1972-04-06
119	Verbena litoralis	Lawai Valley; P.T.B.G		Preserved_Specimen	Herb w/pale blue flowers. Uncommon in valley.	Herbst, D.R.	2296	Kauai	BISH	1972-01-04
120	Verbena litoralis	East Maui, Upper Kaupo Gap	in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground.	Preserved_Specimen	Flowers purple	Higashino, P.K.	885	Maui	BISH	1975-06-22
121	Verbena litoralis	Located on the Mauna Loa Strip road	about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19	Preserved_Specimen	Shrub about 4 ft. tall	Newell, C.L.	966	Hawaii	BISH	1967-06-23
122	Verbena litoralis	near Koko Head, Kaluanui Ridge,Koolau Range	guava zone; at summit	Preserved_Specimen		Egler, F.E.	3783	Oahu	BISH	1937-07-01
123	Verbena litoralis	Kaimuki, Honolulu	roadside	Preserved_Specimen		Fosberg, F.R.	8856	Oahu	BISH	1932-09-24
124	Verbena litoralis	0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline;	on rockpile, vegetation Myoporum-Chamaesyce dominant,	Preserved_Specimen		Wagner, W.L.	4688	Hawaii	BISH	1982-11-17
125	Verbena litoralis	Growing next to Kaluanui Stream	Approximately 1/2 mile from ocean.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-05-18
126	Verbena litoralis	Waianae Mts., Makaha Valley	on trail	Preserved_Specimen		Hasegawa, M.		Oahu	BISH	1932-04-03
127	Verbena litoralis	Keauhou Ranch, Kau District		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/16/2007
128	Verbena litoralis	Kalaoa, HI 96740, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	10/18/2007
129	Verbena litoralis	Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field	Roadside.	Preserved_Specimen		Wagner, W.L.	4826	Oahu	BISH	1983-04-04
130	Verbena litoralis	Honolulu, Hawaii, United States		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
131	Verbena litoralis	Growing next to Kaluanui Stream		OBSERVATION		Curtis Daehler		Oahu	HAPI	3/21/2007
132	Verbena litoralis	S half of island	Ponds	Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
133	Verbena litoralis	East Maui, floor of Haleakala Crater	1/2 mile East of Holua Cabin along trail	Preserved_Specimen		Henrickson, J.S.	3939	Maui	BISH	1969-07-20
134	Verbena litoralis	Maui		Preserved_Specimen	Flower lavender	Handy, E.S.	3320	Maui	BISH	1933-06-10
135	Verbena litoralis	Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ)	Common roadside weed.	Preserved_Specimen		Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
136	Verbena litoralis	ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision	The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica.	Preserved_Specimen		Yoshida, L.	79.053	Hawaii	BISH	1979-02-28
137	Verbena litoralis	Wahiawa		Preserved_Specimen	Erect herb, 1m; flower purple	Swarbrick, J.T.	0.62	Oahu	BISH	1976-01-07
138	Verbena litoralis	Haena		Preserved_Specimen				Kauai	BISH	1953-10-30
139	Verbena litoralis	Weed of pineapple field near northern junction of Highways 37 and 377		Preserved_Specimen	Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple.	Herbst, D.R.	795	Maui	BISH	1967-12-18
140	Verbena litoralis	Kanehoa, Waianaeuka	near top of ridge	Preserved_Specimen		Hume, E.P.	123	Oahu	BISH	1931-03-22
141	Verbena litoralis	East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store	Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus.	Preserved_Specimen		Imada, C.	200292	Maui	BISH	2002-11-19
142	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen	Herb, woody base,1.25 m high	Christophersen, E.		Oahu	BISH	1924-12-07
143	Verbena litoralis	Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility		Preserved_Specimen	Herb 1 m tall; flws light violet.	Lammers, T.G.	5313	Oahu	BISH	1983-08-27
144	Verbena litoralis	Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail		Preserved_Specimen		Wagner, W.L.	5398	Molokai	BISH	1984-07-17
145	Verbena litoralis	Maui		Preserved_Specimen		Forbes, C.N.	2036	Maui	BISH	1920-04-19
146	Verbena litoralis	Hawaii County, HI, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
147	Verbena litoralis	Pohakuloa dry area		Preserved_Specimen		Long, C.R.	4309	Hawaii	BISH	1966-12-23
148	Verbena litoralis	Maui		Preserved_Specimen		Wagner, W.L.	4783	Maui	BISH	1983-01-14
149	Verbena litoralis	Oahu		Preserved_Specimen		Rémy, J.	406	Oahu	BISH	1851-01-01
150	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	141	Lanai	BISH	1913-10-10
151	Verbena litoralis	Area within 300 m west of LZ-1. UTM coordinates 752400-2275900		Preserved_Specimen		Warren, S.	14	Kahoolawe	BISH	1987-04-09
152	Verbena litoralis	Waipio		Preserved_Specimen		Swanson, R.		Hawaii	BISH	1931-08-29
153	Verbena litoralis	Tantalus Rd		Preserved_Specimen		Forbes, A.		Oahu	BISH	1930-07-18
154	Verbena litoralis	Waikolu Valley, head of and E. rim		Preserved_Specimen		Kondo, Y.		Molokai	BISH	1962-06-18
155	Verbena litoralis	Honolulu		Preserved_Specimen		Pukui, M.K.		Oahu	BISH	1931-01-01
156	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen		Christophersen, E.		Oahu	BISH	1924-12-07
157	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
158	Verbena litoralis	Waianae Mts., South end		Preserved_Specimen		Harley, K.		Oahu	BISH	1956-05-04
159	Verbena litoralis	Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea		Preserved_Specimen		Adee, K.T.		Hawaii	BISH
160	Verbena litoralis	Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road		Preserved_Specimen		Ziegler, M.F.		Oahu	BISH	1986-11-08
161	Verbena litoralis	Kealia Salt Marsh		Preserved_Specimen		Funk, E.J.		Maui	BISH	1990-02-15
162	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
163	Verbena litoralis	Mokuleia Forest Reserve, Waianae Mts		Preserved_Specimen		Onouye, R.		Oahu	BISH	1934-02-04
164	Verbena litoralis	Valley above country club, Honolulu		Preserved_Specimen		Forbes, C.N.		Oahu	BISH	1908-07-08
165	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
166	Verbena litoralis	Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above		Preserved_Specimen		Sohmer, S.H.	5995	Oahu	BISH	1969-08-27
167	Verbena litoralis	Lanai		Preserved_Specimen		Munro, G.C.	292	Lanai	BISH
168	Verbena litoralis	Slopes of Mauna Kea (Wailei)		Preserved_Specimen		Forbes, C.N.	465	Hawaii	BISH	1911-08-01
169	Verbena litoralis	Kaua Trail, Waianae Mts		Preserved_Specimen		Woolford, H.A.	200	Oahu	BISH	1955-12-11
170	Verbena litoralis	Kealia trail		Preserved_Specimen		Bennett, T.	244	Oahu	BISH	1982-03-25
171	Verbena litoralis	Kualoa Pasture		Preserved_Specimen		Rock, J.F.C.	3007	Oahu	BISH	1942-01-01
172	Verbena litoralis	Kaholuamanoa, above Waimea		Preserved_Specimen		Heller, A.A.	2046	Kauai	BISH	1895-09-02
173	Verbena litoralis	Pauoa Valley		Preserved_Specimen		Garber, D.W.	339	Oahu	BISH	1920-03-21
174	Verbena litoralis	Honolulu, Aihualama		Preserved_Specimen		Russell, P.	253	Oahu	BISH	1932-07-08
175	Verbena litoralis	Crater of Haleakala		Preserved_Specimen		Forbes, C.N.	1083	Maui	BISH	1919-08-09
176	Verbena litoralis	Koele		Preserved_Specimen		Forbes, C.N.	188	Lanai	BISH	1918-06-01
177	Verbena litoralis	Maui		Preserved_Specimen		Neal, M.C.		Maui	BISH	1933-08-01
178	Verbena litoralis	unknown		Preserved_Specimen		Mann, H.	437		BISH	1864-01-01
179	Verbena litoralis	Kilauea, Kaloko Reservoir		Preserved_Specimen		Forbes, C.N.	572	Kauai	BISH	1916-10-11
180	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	365	Lanai	BISH	1928-08-01
181	Verbena litoralis	Palolo Valley, middle ridge	forest	Preserved_Specimen		Christophersen, E.	1666	Oahu	BISH	1931-03-26
182	Verbena litoralis	Palaau, west side of airport	pineapple field; dry, exposed, windy.	Preserved_Specimen	Herb to sub-shrub with blue flowers	Herbst, D.R.	5092	Molokai	BISH	1974-11-11
183	Verbena litoralis	Mt. Kalena, Waianae	Lantana grove	Preserved_Specimen	herb 4 ft.	Grant, M.L.	7445	Oahu	BISH	1934-09-30
184	Verbena litoralis	Koloa, Kalaheo, upper end of Puuwai Rd.	wet roadside secondary forest; Wedelia	Preserved_Specimen	to 0.5 m herb, slender; square stem, small purple flowers	Trauernicht, C.	205	Kauai	BISH	2007-10-15
185	Verbena litoralis	Nawaimaka drainage.	Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai	Preserved_Specimen	1.5 m. tall, stems green, corolla pale pruple.	Wood, K.R.	13816	Kauai	BISH	2009-07-16
186	Verbena litoralis	ridge north of Kuapa Pond	open, hillsides	Preserved_Specimen		Gillespie, J.W.	1076	Oahu	BISH	1927-07-08
187	Verbena litoralis	Kau District: Keauhou Ranch	Along fenceline of horse corrall in lower pastures of ranch.	Preserved_Specimen	Herb, 3 ft. tall, purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
188	Verbena litoralis	stream bank below Dillingham Field wetland	grasses	Preserved_Specimen		Funk, E.J.	1105	Oahu	BISH	1990-05-08
189	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
190	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
191	Verbena litoralis	Makanikahau Reservoir, Old	Dried mud	Preserved_Specimen		St.John, H.	23633	Niihau	BISH	1949-04-01
192	Verbena litoralis	Puu Huluhulu. Near the escarpment Transect (T16)	Remnant dryland community. Common subshrub among Pennisetum setaceum.	Preserved_Specimen	Common subshrub.	Takeuchi, W.N.	5714	Hawaii	BISH	1989-12-26
193	Verbena litoralis	Kaluanui Flats	Pineapple fields	Preserved_Specimen	Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas.	Spence, G.	369	Lanai	BISH	1973-10-24
194	Verbena litoralis	Mauna Kea (Puu Laau Road)	In open Mamane-Naio forest.	Preserved_Specimen		Engilis Jr., A.	39	Hawaii	BISH	1987-08-19
195	Verbena litoralis	Keanakolu, Hilo Forest Reserve	Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus.	Preserved_Specimen		Higashino, P.K.	6965	Hawaii	BISH	1978-03-01
196	Verbena litoralis	Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost	in open sunny area.	Preserved_Specimen		Herat, T.	559	Hawaii	BISH	1974-06-12
197	Verbena litoralis	North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary	Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa	Preserved_Specimen		Close, B.	109426	Hawaii	BISH	1994-10-29
198	Verbena litoralis	Peacock Flat, Waianae Mts	roadside, dry forest	Preserved_Specimen	herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue.	Spence, G.	209	Oahu	BISH	1973-09-06
199	Verbena litoralis	Puu Laau	Weed growing near the hunter's cabin.	Preserved_Specimen		Herbst, D.R.	5210	Hawaii	BISH	1975-01-18
200	Verbena litoralis	Kahaluu; Kailua; Kapua	Koa forest.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
201	Verbena litoralis	Puuwaawaa; track to 1986 burn area (from Ranch HQ)	4WD access track to burn area.	Preserved_Specimen	Common roadside weed.	Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
202	Verbena litoralis	Kawa side of Kahana Valley, pass the main gate and the marshy area	along jeep trail, open areas	Preserved_Specimen	common weed, ~4', with light blue flowers	Herat, T.	158	Oahu	BISH	1972-08-14
203	Verbena litoralis	1750' Field 123, Honokaa GSugar Co. Hamakua District	Common weed at field edge, and on jeep roads through field, not present inside mature stand.	Preserved_Specimen	Perennial root, plants 30- 48 tall, corolla pale blue	Tomich, P.Q.	911	Hawaii	BISH	1960-03-18
204	Verbena litoralis	Kahuku, Ocean View Estates,	open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive.	Preserved_Specimen	Common roadside weed with blue flowers.	Herbst, D.R.	5068	Hawaii	BISH	1974-10-09
205	Verbena litoralis	Kilauea	weed in abandoned sorghum field	Preserved_Specimen		Herbst, D.R.	5244	Kauai	BISH	1975-02-27
206	Verbena litoralis	Laie	clay, pasture	Preserved_Specimen	herb 1-6', tall slender stems	Guinto, J.	27	Oahu	BISH	1986-03-06
207	Verbena litoralis	Honolulu, 3830 Tantalus Dr., access road to Puu Ohia	roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum.	Preserved_Specimen	to 4-20 dm tall; corolla blue	Ray, G.	50	Oahu	BISH	1997-09-15
208	Verbena litoralis	Kewalo, back of Punchbowl	open, dry, roadside	Preserved_Specimen	1 m tall; flower - small, inconspicous, lavender	Judd, C.S.	11	Oahu	BISH	1924-11-06
209	Verbena litoralis	Lower half of Kaala Road	mesic forest	Preserved_Specimen	flowers lavender	Annable, C.R.	3727	Oahu	BISH	1997-10-15
210	Verbena litoralis	E central part of island, slope of Moaula	Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping.	Preserved_Specimen	Flowers purple.	Cuddihy, L.W.	428	Kahoolawe	BISH	1980-04-25
211	Verbena litoralis	Waikane-Schofield trail, Koolau Mts, Kahana	along trail	Preserved_Specimen	fls. lavendar	Fosberg, F.R.	10276	Oahu	BISH	1933-09-22
212	Verbena litoralis	Trail to Kaluiwaa (Sacred Falls), outside forest Reserve	At base of foot path trail.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-03-21
213	Verbena litoralis	East Maui, Haleakala Crater, Kaupo Gap	Dry volcanic gravel by trail.	Preserved_Specimen	Flws lavender-blue.	St.John, H.	21182	Maui	BISH	1945-08-27
214	Verbena litoralis	Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road	Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside..	Preserved_Specimen		Newell, C.L.	471	Hawaii	BISH	1966-08-20
215	Verbena litoralis	Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979	Pohakuloa burn 1979.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
216	Verbena litoralis	ahupuaa of Halepuaa Forest Reserve in the Puna District	Found growing on the side of the old railroad.	Preserved_Specimen		ESP Field Crew		Hawaii	BISH	1979-08-15
217	Verbena litoralis	North Kohala District, Kalala ahupua'a, Lahikiola cinder cone	In Kikuyu grassland	Preserved_Specimen		Bunch, P.M.	112	Hawaii	BISH	1989-03-13
218	Verbena litoralis	Kipapa Gulch, Wahiawa	moderately moist pineapple field	Preserved_Specimen		Hosaka, E.Y.	422	Oahu	BISH	1931-06-18
219	Verbena litoralis	Waimea Audubon Center, Waimea Valley, Oahu, Hawaii	Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions.	Preserved_Specimen	Shrub, spindly, flowers purple, on spikes.	Crago, L.M.	2005207	Oahu	BISH	2005-10-18
220	Verbena litoralis	Waiapaa Gulch, flat area near the road above	Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants.	Preserved_Specimen		Funk, E.J.	920004	Lanai	BISH	1992-01-26
221	Verbena litoralis	South Kohala: Waimea, Lalamilo	Range paddock; occasional.	Preserved_Specimen	Branching plant 1-3 ft.	Hosaka, E.Y.	2122	Hawaii	BISH	1938-06-01
222	Verbena litoralis	Kaalualu Bay near South Point	Only one individual found in waste area.	Preserved_Specimen	Herb up to one m in height with small purplish flowers.	Whistler, W.A.	42	Hawaii	BISH	1972-04-06
223	Verbena litoralis	Lawai Valley; P.T.B.G		Preserved_Specimen	Herb w/pale blue flowers. Uncommon in valley.	Herbst, D.R.	2296	Kauai	BISH	1972-01-04
224	Verbena litoralis	East Maui, Upper Kaupo Gap	in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground.	Preserved_Specimen	Flowers purple	Higashino, P.K.	885	Maui	BISH	1975-06-22
225	Verbena litoralis	Located on the Mauna Loa Strip road	about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19	Preserved_Specimen	Shrub about 4 ft. tall	Newell, C.L.	966	Hawaii	BISH	1967-06-23
226	Verbena litoralis	near Koko Head, Kaluanui Ridge,Koolau Range	guava zone; at summit	Preserved_Specimen		Egler, F.E.	3783	Oahu	BISH	1937-07-01
227	Verbena litoralis	Kaimuki, Honolulu	roadside	Preserved_Specimen		Fosberg, F.R.	8856	Oahu	BISH	1932-09-24
228	Verbena litoralis	0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline;	on rockpile, vegetation Myoporum-Chamaesyce dominant,	Preserved_Specimen		Wagner, W.L.	4688	Hawaii	BISH	1982-11-17
229	Verbena litoralis	Growing next to Kaluanui Stream	Approximately 1/2 mile from ocean.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-05-18
230	Verbena litoralis	Waianae Mts., Makaha Valley	on trail	Preserved_Specimen		Hasegawa, M.		Oahu	BISH	1932-04-03
231	Verbena litoralis	Keauhou Ranch, Kau District		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/16/2007
232	Verbena litoralis	Kalaoa, HI 96740, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	10/18/2007
233	Verbena litoralis	Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field	Roadside.	Preserved_Specimen		Wagner, W.L.	4826	Oahu	BISH	1983-04-04
234	Verbena litoralis	Honolulu, Hawaii, United States		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
235	Verbena litoralis	Growing next to Kaluanui Stream		OBSERVATION		Curtis Daehler		Oahu	HAPI	3/21/2007
236	Verbena litoralis	S half of island	Ponds	Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
237	Verbena litoralis	East Maui, floor of Haleakala Crater	1/2 mile East of Holua Cabin along trail	Preserved_Specimen		Henrickson, J.S.	3939	Maui	BISH	1969-07-20
238	Verbena litoralis	Maui		Preserved_Specimen	Flower lavender	Handy, E.S.	3320	Maui	BISH	1933-06-10
239	Verbena litoralis	Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ)	Common roadside weed.	Preserved_Specimen		Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
240	Verbena litoralis	ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision	The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica.	Preserved_Specimen		Yoshida, L.	79.053	Hawaii	BISH	1979-02-28
241	Verbena litoralis	Wahiawa		Preserved_Specimen	Erect herb, 1m; flower purple	Swarbrick, J.T.	0.62	Oahu	BISH	1976-01-07
242	Verbena litoralis	Haena		Preserved_Specimen				Kauai	BISH	1953-10-30
243	Verbena litoralis	Weed of pineapple field near northern junction of Highways 37 and 377		Preserved_Specimen	Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple.	Herbst, D.R.	795	Maui	BISH	1967-12-18
244	Verbena litoralis	Kanehoa, Waianaeuka	near top of ridge	Preserved_Specimen		Hume, E.P.	123	Oahu	BISH	1931-03-22
245	Verbena litoralis	East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store	Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus.	Preserved_Specimen		Imada, C.	200292	Maui	BISH	2002-11-19
246	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen	Herb, woody base,1.25 m high	Christophersen, E.		Oahu	BISH	1924-12-07
247	Verbena litoralis	Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility		Preserved_Specimen	Herb 1 m tall; flws light violet.	Lammers, T.G.	5313	Oahu	BISH	1983-08-27
248	Verbena litoralis	Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail		Preserved_Specimen		Wagner, W.L.	5398	Molokai	BISH	1984-07-17
249	Verbena litoralis	Maui		Preserved_Specimen		Forbes, C.N.	2036	Maui	BISH	1920-04-19
250	Verbena litoralis	Hawaii County, HI, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
251	Verbena litoralis	Pohakuloa dry area		Preserved_Specimen		Long, C.R.	4309	Hawaii	BISH	1966-12-23
252	Verbena litoralis	Maui		Preserved_Specimen		Wagner, W.L.	4783	Maui	BISH	1983-01-14
253	Verbena litoralis	Oahu		Preserved_Specimen		Rémy, J.	406	Oahu	BISH	1851-01-01
254	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	141	Lanai	BISH	1913-10-10
255	Verbena litoralis	Area within 300 m west of LZ-1. UTM coordinates 752400-2275900		Preserved_Specimen		Warren, S.	14	Kahoolawe	BISH	1987-04-09
256	Verbena litoralis	Waipio		Preserved_Specimen		Swanson, R.		Hawaii	BISH	1931-08-29
257	Verbena litoralis	Tantalus Rd		Preserved_Specimen		Forbes, A.		Oahu	BISH	1930-07-18
258	Verbena litoralis	Waikolu Valley, head of and E. rim		Preserved_Specimen		Kondo, Y.		Molokai	BISH	1962-06-18
259	Verbena litoralis	Honolulu		Preserved_Specimen		Pukui, M.K.		Oahu	BISH	1931-01-01
260	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen		Christophersen, E.		Oahu	BISH	1924-12-07
261	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
262	Verbena litoralis	Waianae Mts., South end		Preserved_Specimen		Harley, K.		Oahu	BISH	1956-05-04
263	Verbena litoralis	Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea		Preserved_Specimen		Adee, K.T.		Hawaii	BISH
264	Verbena litoralis	Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road		Preserved_Specimen		Ziegler, M.F.		Oahu	BISH	1986-11-08
265	Verbena litoralis	Kealia Salt Marsh		Preserved_Specimen		Funk, E.J.		Maui	BISH	1990-02-15
266	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
267	Verbena litoralis	Mokuleia Forest Reserve, Waianae Mts		Preserved_Specimen		Onouye, R.		Oahu	BISH	1934-02-04
268	Verbena litoralis	Valley above country club, Honolulu		Preserved_Specimen		Forbes, C.N.		Oahu	BISH	1908-07-08
269	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
270	Verbena litoralis	Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above		Preserved_Specimen		Sohmer, S.H.	5995	Oahu	BISH	1969-08-27
271	Verbena litoralis	Lanai		Preserved_Specimen		Munro, G.C.	292	Lanai	BISH
272	Verbena litoralis	Slopes of Mauna Kea (Wailei)		Preserved_Specimen		Forbes, C.N.	465	Hawaii	BISH	1911-08-01
273	Verbena litoralis	Kaua Trail, Waianae Mts		Preserved_Specimen		Woolford, H.A.	200	Oahu	BISH	1955-12-11
274	Verbena litoralis	Kealia trail		Preserved_Specimen		Bennett, T.	244	Oahu	BISH	1982-03-25
275	Verbena litoralis	Kualoa Pasture		Preserved_Specimen		Rock, J.F.C.	3007	Oahu	BISH	1942-01-01
276	Verbena litoralis	Kaholuamanoa, above Waimea		Preserved_Specimen		Heller, A.A.	2046	Kauai	BISH	1895-09-02
277	Verbena litoralis	Pauoa Valley		Preserved_Specimen		Garber, D.W.	339	Oahu	BISH	1920-03-21
278	Verbena litoralis	Honolulu, Aihualama		Preserved_Specimen		Russell, P.	253	Oahu	BISH	1932-07-08
279	Verbena litoralis	Crater of Haleakala		Preserved_Specimen		Forbes, C.N.	1083	Maui	BISH	1919-08-09
280	Verbena litoralis	Koele		Preserved_Specimen		Forbes, C.N.	188	Lanai	BISH	1918-06-01
281	Verbena litoralis	Maui		Preserved_Specimen		Neal, M.C.		Maui	BISH	1933-08-01
282	Verbena litoralis	unknown		Preserved_Specimen		Mann, H.	437		BISH	1864-01-01
283	Verbena litoralis	Kilauea, Kaloko Reservoir		Preserved_Specimen		Forbes, C.N.	572	Kauai	BISH	1916-10-11
284	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	365	Lanai	BISH	1928-08-01
285	Verbena litoralis	Palolo Valley, middle ridge	forest	Preserved_Specimen		Christophersen, E.	1666	Oahu	BISH	1931-03-26
286	Verbena litoralis	Palaau, west side of airport	pineapple field; dry, exposed, windy.	Preserved_Specimen	Herb to sub-shrub with blue flowers	Herbst, D.R.	5092	Molokai	BISH	1974-11-11
287	Verbena litoralis	Mt. Kalena, Waianae	Lantana grove	Preserved_Specimen	herb 4 ft.	Grant, M.L.	7445	Oahu	BISH	1934-09-30
288	Verbena litoralis	Koloa, Kalaheo, upper end of Puuwai Rd.	wet roadside secondary forest; Wedelia	Preserved_Specimen	to 0.5 m herb, slender; square stem, small purple flowers	Trauernicht, C.	205	Kauai	BISH	2007-10-15
289	Verbena litoralis	Nawaimaka drainage.	Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai	Preserved_Specimen	1.5 m. tall, stems green, corolla pale pruple.	Wood, K.R.	13816	Kauai	BISH	2009-07-16
290	Verbena litoralis	ridge north of Kuapa Pond	open, hillsides	Preserved_Specimen		Gillespie, J.W.	1076	Oahu	BISH	1927-07-08
291	Verbena litoralis	Kau District: Keauhou Ranch	Along fenceline of horse corrall in lower pastures of ranch.	Preserved_Specimen	Herb, 3 ft. tall, purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
292	Verbena litoralis	stream bank below Dillingham Field wetland	grasses	Preserved_Specimen		Funk, E.J.	1105	Oahu	BISH	1990-05-08
293	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
294	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
295	Verbena litoralis	Makanikahau Reservoir, Old	Dried mud	Preserved_Specimen		St.John, H.	23633	Niihau	BISH	1949-04-01
296	Verbena litoralis	Puu Huluhulu. Near the escarpment Transect (T16)	Remnant dryland community. Common subshrub among Pennisetum setaceum.	Preserved_Specimen	Common subshrub.	Takeuchi, W.N.	5714	Hawaii	BISH	1989-12-26
297	Verbena litoralis	Kaluanui Flats	Pineapple fields	Preserved_Specimen	Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas.	Spence, G.	369	Lanai	BISH	1973-10-24
298	Verbena litoralis	Mauna Kea (Puu Laau Road)	In open Mamane-Naio forest.	Preserved_Specimen		Engilis Jr., A.	39	Hawaii	BISH	1987-08-19
299	Verbena litoralis	Keanakolu, Hilo Forest Reserve	Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus.	Preserved_Specimen		Higashino, P.K.	6965	Hawaii	BISH	1978-03-01
300	Verbena litoralis	Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost	in open sunny area.	Preserved_Specimen		Herat, T.	559	Hawaii	BISH	1974-06-12
301	Verbena litoralis	North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary	Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa	Preserved_Specimen		Close, B.	109426	Hawaii	BISH	1994-10-29
302	Verbena litoralis	Peacock Flat, Waianae Mts	roadside, dry forest	Preserved_Specimen	herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue.	Spence, G.	209	Oahu	BISH	1973-09-06
303	Verbena litoralis	Puu Laau	Weed growing near the hunter's cabin.	Preserved_Specimen		Herbst, D.R.	5210	Hawaii	BISH	1975-01-18
304	Verbena litoralis	Kahaluu; Kailua; Kapua	Koa forest.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
305	Verbena litoralis	Puuwaawaa; track to 1986 burn area (from Ranch HQ)	4WD access track to burn area.	Preserved_Specimen	Common roadside weed.	Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
306	Verbena litoralis	Kawa side of Kahana Valley, pass the main gate and the marshy area	along jeep trail, open areas	Preserved_Specimen	common weed, ~4', with light blue flowers	Herat, T.	158	Oahu	BISH	1972-08-14
307	Verbena litoralis	1750' Field 123, Honokaa GSugar Co. Hamakua District	Common weed at field edge, and on jeep roads through field, not present inside mature stand.	Preserved_Specimen	Perennial root, plants 30- 48 tall, corolla pale blue	Tomich, P.Q.	911	Hawaii	BISH	1960-03-18
308	Verbena litoralis	Kahuku, Ocean View Estates,	open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive.	Preserved_Specimen	Common roadside weed with blue flowers.	Herbst, D.R.	5068	Hawaii	BISH	1974-10-09
309	Verbena litoralis	Kilauea	weed in abandoned sorghum field	Preserved_Specimen		Herbst, D.R.	5244	Kauai	BISH	1975-02-27
310	Verbena litoralis	Laie	clay, pasture	Preserved_Specimen	herb 1-6', tall slender stems	Guinto, J.	27	Oahu	BISH	1986-03-06
311	Verbena litoralis	Honolulu, 3830 Tantalus Dr., access road to Puu Ohia	roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum.	Preserved_Specimen	to 4-20 dm tall; corolla blue	Ray, G.	50	Oahu	BISH	1997-09-15
312	Verbena litoralis	Kewalo, back of Punchbowl	open, dry, roadside	Preserved_Specimen	1 m tall; flower - small, inconspicous, lavender	Judd, C.S.	11	Oahu	BISH	1924-11-06
313	Verbena litoralis	Lower half of Kaala Road	mesic forest	Preserved_Specimen	flowers lavender	Annable, C.R.	3727	Oahu	BISH	1997-10-15
314	Verbena litoralis	E central part of island, slope of Moaula	Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping.	Preserved_Specimen	Flowers purple.	Cuddihy, L.W.	428	Kahoolawe	BISH	1980-04-25
315	Verbena litoralis	Waikane-Schofield trail, Koolau Mts, Kahana	along trail	Preserved_Specimen	fls. lavendar	Fosberg, F.R.	10276	Oahu	BISH	1933-09-22
316	Verbena litoralis	Trail to Kaluiwaa (Sacred Falls), outside forest Reserve	At base of foot path trail.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-03-21
317	Verbena litoralis	East Maui, Haleakala Crater, Kaupo Gap	Dry volcanic gravel by trail.	Preserved_Specimen	Flws lavender-blue.	St.John, H.	21182	Maui	BISH	1945-08-27
318	Verbena litoralis	Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road	Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside..	Preserved_Specimen		Newell, C.L.	471	Hawaii	BISH	1966-08-20
319	Verbena litoralis	Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979	Pohakuloa burn 1979.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
320	Verbena litoralis	ahupuaa of Halepuaa Forest Reserve in the Puna District	Found growing on the side of the old railroad.	Preserved_Specimen		ESP Field Crew		Hawaii	BISH	1979-08-15
321	Verbena litoralis	North Kohala District, Kalala ahupua'a, Lahikiola cinder cone	In Kikuyu grassland	Preserved_Specimen		Bunch, P.M.	112	Hawaii	BISH	1989-03-13
322	Verbena litoralis	Kipapa Gulch, Wahiawa	moderately moist pineapple field	Preserved_Specimen		Hosaka, E.Y.	422	Oahu	BISH	1931-06-18
323	Verbena litoralis	Waimea Audubon Center, Waimea Valley, Oahu, Hawaii	Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions.	Preserved_Specimen	Shrub, spindly, flowers purple, on spikes.	Crago, L.M.	2005207	Oahu	BISH	2005-10-18
324	Verbena litoralis	Waiapaa Gulch, flat area near the road above	Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants.	Preserved_Specimen		Funk, E.J.	920004	Lanai	BISH	1992-01-26
325	Verbena litoralis	South Kohala: Waimea, Lalamilo	Range paddock; occasional.	Preserved_Specimen	Branching plant 1-3 ft.	Hosaka, E.Y.	2122	Hawaii	BISH	1938-06-01
326	Verbena litoralis	Kaalualu Bay near South Point	Only one individual found in waste area.	Preserved_Specimen	Herb up to one m in height with small purplish flowers.	Whistler, W.A.	42	Hawaii	BISH	1972-04-06
327	Verbena litoralis	Lawai Valley; P.T.B.G		Preserved_Specimen	Herb w/pale blue flowers. Uncommon in valley.	Herbst, D.R.	2296	Kauai	BISH	1972-01-04
328	Verbena litoralis	East Maui, Upper Kaupo Gap	in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground.	Preserved_Specimen	Flowers purple	Higashino, P.K.	885	Maui	BISH	1975-06-22
329	Verbena litoralis	Located on the Mauna Loa Strip road	about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19	Preserved_Specimen	Shrub about 4 ft. tall	Newell, C.L.	966	Hawaii	BISH	1967-06-23
330	Verbena litoralis	near Koko Head, Kaluanui Ridge,Koolau Range	guava zone; at summit	Preserved_Specimen		Egler, F.E.	3783	Oahu	BISH	1937-07-01
331	Verbena litoralis	Kaimuki, Honolulu	roadside	Preserved_Specimen		Fosberg, F.R.	8856	Oahu	BISH	1932-09-24
332	Verbena litoralis	0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline;	on rockpile, vegetation Myoporum-Chamaesyce dominant,	Preserved_Specimen		Wagner, W.L.	4688	Hawaii	BISH	1982-11-17
333	Verbena litoralis	Growing next to Kaluanui Stream	Approximately 1/2 mile from ocean.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-05-18
334	Verbena litoralis	Waianae Mts., Makaha Valley	on trail	Preserved_Specimen		Hasegawa, M.		Oahu	BISH	1932-04-03
335	Verbena litoralis	Keauhou Ranch, Kau District		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/16/2007
336	Verbena litoralis	Kalaoa, HI 96740, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	10/18/2007
337	Verbena litoralis	Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field	Roadside.	Preserved_Specimen		Wagner, W.L.	4826	Oahu	BISH	1983-04-04
338	Verbena litoralis	Honolulu, Hawaii, United States		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
339	Verbena litoralis	Growing next to Kaluanui Stream		OBSERVATION		Curtis Daehler		Oahu	HAPI	3/21/2007
340	Verbena litoralis	S half of island	Ponds	Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
341	Verbena litoralis	East Maui, floor of Haleakala Crater	1/2 mile East of Holua Cabin along trail	Preserved_Specimen		Henrickson, J.S.	3939	Maui	BISH	1969-07-20
342	Verbena litoralis	Maui		Preserved_Specimen	Flower lavender	Handy, E.S.	3320	Maui	BISH	1933-06-10
343	Verbena litoralis	Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ)	Common roadside weed.	Preserved_Specimen		Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
344	Verbena litoralis	ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision	The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica.	Preserved_Specimen		Yoshida, L.	79.053	Hawaii	BISH	1979-02-28
345	Verbena litoralis	Wahiawa		Preserved_Specimen	Erect herb, 1m; flower purple	Swarbrick, J.T.	0.62	Oahu	BISH	1976-01-07
346	Verbena litoralis	Haena		Preserved_Specimen				Kauai	BISH	1953-10-30
347	Verbena litoralis	Weed of pineapple field near northern junction of Highways 37 and 377		Preserved_Specimen	Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple.	Herbst, D.R.	795	Maui	BISH	1967-12-18
348	Verbena litoralis	Kanehoa, Waianaeuka	near top of ridge	Preserved_Specimen		Hume, E.P.	123	Oahu	BISH	1931-03-22
349	Verbena litoralis	East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store	Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus.	Preserved_Specimen		Imada, C.	200292	Maui	BISH	2002-11-19
350	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen	Herb, woody base,1.25 m high	Christophersen, E.		Oahu	BISH	1924-12-07
351	Verbena litoralis	Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility		Preserved_Specimen	Herb 1 m tall; flws light violet.	Lammers, T.G.	5313	Oahu	BISH	1983-08-27
352	Verbena litoralis	Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail		Preserved_Specimen		Wagner, W.L.	5398	Molokai	BISH	1984-07-17
353	Verbena litoralis	Maui		Preserved_Specimen		Forbes, C.N.	2036	Maui	BISH	1920-04-19
354	Verbena litoralis	Hawaii County, HI, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
355	Verbena litoralis	Pohakuloa dry area		Preserved_Specimen		Long, C.R.	4309	Hawaii	BISH	1966-12-23
356	Verbena litoralis	Maui		Preserved_Specimen		Wagner, W.L.	4783	Maui	BISH	1983-01-14
357	Verbena litoralis	Oahu		Preserved_Specimen		Rémy, J.	406	Oahu	BISH	1851-01-01
358	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	141	Lanai	BISH	1913-10-10
359	Verbena litoralis	Area within 300 m west of LZ-1. UTM coordinates 752400-2275900		Preserved_Specimen		Warren, S.	14	Kahoolawe	BISH	1987-04-09
360	Verbena litoralis	Waipio		Preserved_Specimen		Swanson, R.		Hawaii	BISH	1931-08-29
361	Verbena litoralis	Tantalus Rd		Preserved_Specimen		Forbes, A.		Oahu	BISH	1930-07-18
362	Verbena litoralis	Waikolu Valley, head of and E. rim		Preserved_Specimen		Kondo, Y.		Molokai	BISH	1962-06-18
363	Verbena litoralis	Honolulu		Preserved_Specimen		Pukui, M.K.		Oahu	BISH	1931-01-01
364	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen		Christophersen, E.		Oahu	BISH	1924-12-07
365	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
366	Verbena litoralis	Waianae Mts., South end		Preserved_Specimen		Harley, K.		Oahu	BISH	1956-05-04
367	Verbena litoralis	Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea		Preserved_Specimen		Adee, K.T.		Hawaii	BISH
368	Verbena litoralis	Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road		Preserved_Specimen		Ziegler, M.F.		Oahu	BISH	1986-11-08
369	Verbena litoralis	Kealia Salt Marsh		Preserved_Specimen		Funk, E.J.		Maui	BISH	1990-02-15
370	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
371	Verbena litoralis	Mokuleia Forest Reserve, Waianae Mts		Preserved_Specimen		Onouye, R.		Oahu	BISH	1934-02-04
372	Verbena litoralis	Valley above country club, Honolulu		Preserved_Specimen		Forbes, C.N.		Oahu	BISH	1908-07-08
373	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
374	Verbena litoralis	Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above		Preserved_Specimen		Sohmer, S.H.	5995	Oahu	BISH	1969-08-27
375	Verbena litoralis	Lanai		Preserved_Specimen		Munro, G.C.	292	Lanai	BISH
376	Verbena litoralis	Slopes of Mauna Kea (Wailei)		Preserved_Specimen		Forbes, C.N.	465	Hawaii	BISH	1911-08-01
377	Verbena litoralis	Kaua Trail, Waianae Mts		Preserved_Specimen		Woolford, H.A.	200	Oahu	BISH	1955-12-11
378	Verbena litoralis	Kealia trail		Preserved_Specimen		Bennett, T.	244	Oahu	BISH	1982-03-25
379	Verbena litoralis	Kualoa Pasture		Preserved_Specimen		Rock, J.F.C.	3007	Oahu	BISH	1942-01-01
380	Verbena litoralis	Kaholuamanoa, above Waimea		Preserved_Specimen		Heller, A.A.	2046	Kauai	BISH	1895-09-02
381	Verbena litoralis	Pauoa Valley		Preserved_Specimen		Garber, D.W.	339	Oahu	BISH	1920-03-21
382	Verbena litoralis	Honolulu, Aihualama		Preserved_Specimen		Russell, P.	253	Oahu	BISH	1932-07-08
383	Verbena litoralis	Crater of Haleakala		Preserved_Specimen		Forbes, C.N.	1083	Maui	BISH	1919-08-09
384	Verbena litoralis	Koele		Preserved_Specimen		Forbes, C.N.	188	Lanai	BISH	1918-06-01
385	Verbena litoralis	Maui		Preserved_Specimen		Neal, M.C.		Maui	BISH	1933-08-01
386	Verbena litoralis	unknown		Preserved_Specimen		Mann, H.	437		BISH	1864-01-01
387	Verbena litoralis	Kilauea, Kaloko Reservoir		Preserved_Specimen		Forbes, C.N.	572	Kauai	BISH	1916-10-11
388	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	365	Lanai	BISH	1928-08-01
389	Verbena litoralis	Palolo Valley, middle ridge	forest	Preserved_Specimen		Christophersen, E.	1666	Oahu	BISH	1931-03-26
390	Verbena litoralis	Palaau, west side of airport	pineapple field; dry, exposed, windy.	Preserved_Specimen	Herb to sub-shrub with blue flowers	Herbst, D.R.	5092	Molokai	BISH	1974-11-11
391	Verbena litoralis	Mt. Kalena, Waianae	Lantana grove	Preserved_Specimen	herb 4 ft.	Grant, M.L.	7445	Oahu	BISH	1934-09-30
392	Verbena litoralis	Koloa, Kalaheo, upper end of Puuwai Rd.	wet roadside secondary forest; Wedelia	Preserved_Specimen	to 0.5 m herb, slender; square stem, small purple flowers	Trauernicht, C.	205	Kauai	BISH	2007-10-15
393	Verbena litoralis	Nawaimaka drainage.	Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai	Preserved_Specimen	1.5 m. tall, stems green, corolla pale pruple.	Wood, K.R.	13816	Kauai	BISH	2009-07-16
394	Verbena litoralis	ridge north of Kuapa Pond	open, hillsides	Preserved_Specimen		Gillespie, J.W.	1076	Oahu	BISH	1927-07-08
395	Verbena litoralis	Kau District: Keauhou Ranch	Along fenceline of horse corrall in lower pastures of ranch.	Preserved_Specimen	Herb, 3 ft. tall, purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
396	Verbena litoralis	stream bank below Dillingham Field wetland	grasses	Preserved_Specimen		Funk, E.J.	1105	Oahu	BISH	1990-05-08
397	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
398	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
399	Verbena litoralis	Makanikahau Reservoir, Old	Dried mud	Preserved_Specimen		St.John, H.	23633	Niihau	BISH	1949-04-01
400	Verbena litoralis	Puu Huluhulu. Near the escarpment Transect (T16)	Remnant dryland community. Common subshrub among Pennisetum setaceum.	Preserved_Specimen	Common subshrub.	Takeuchi, W.N.	5714	Hawaii	BISH	1989-12-26
401	Verbena litoralis	Kaluanui Flats	Pineapple fields	Preserved_Specimen	Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas.	Spence, G.	369	Lanai	BISH	1973-10-24
402	Verbena litoralis	Mauna Kea (Puu Laau Road)	In open Mamane-Naio forest.	Preserved_Specimen		Engilis Jr., A.	39	Hawaii	BISH	1987-08-19
403	Verbena litoralis	Keanakolu, Hilo Forest Reserve	Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus.	Preserved_Specimen		Higashino, P.K.	6965	Hawaii	BISH	1978-03-01
404	Verbena litoralis	Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost	in open sunny area.	Preserved_Specimen		Herat, T.	559	Hawaii	BISH	1974-06-12
405	Verbena litoralis	North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary	Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa	Preserved_Specimen		Close, B.	109426	Hawaii	BISH	1994-10-29
406	Verbena litoralis	Peacock Flat, Waianae Mts	roadside, dry forest	Preserved_Specimen	herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue.	Spence, G.	209	Oahu	BISH	1973-09-06
407	Verbena litoralis	Puu Laau	Weed growing near the hunter's cabin.	Preserved_Specimen		Herbst, D.R.	5210	Hawaii	BISH	1975-01-18
408	Verbena litoralis	Kahaluu; Kailua; Kapua	Koa forest.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
409	Verbena litoralis	Puuwaawaa; track to 1986 burn area (from Ranch HQ)	4WD access track to burn area.	Preserved_Specimen	Common roadside weed.	Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
410	Verbena litoralis	Kawa side of Kahana Valley, pass the main gate and the marshy area	along jeep trail, open areas	Preserved_Specimen	common weed, ~4', with light blue flowers	Herat, T.	158	Oahu	BISH	1972-08-14
411	Verbena litoralis	1750' Field 123, Honokaa GSugar Co. Hamakua District	Common weed at field edge, and on jeep roads through field, not present inside mature stand.	Preserved_Specimen	Perennial root, plants 30- 48 tall, corolla pale blue	Tomich, P.Q.	911	Hawaii	BISH	1960-03-18
412	Verbena litoralis	Kahuku, Ocean View Estates,	open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive.	Preserved_Specimen	Common roadside weed with blue flowers.	Herbst, D.R.	5068	Hawaii	BISH	1974-10-09
413	Verbena litoralis	Kilauea	weed in abandoned sorghum field	Preserved_Specimen		Herbst, D.R.	5244	Kauai	BISH	1975-02-27
414	Verbena litoralis	Laie	clay, pasture	Preserved_Specimen	herb 1-6', tall slender stems	Guinto, J.	27	Oahu	BISH	1986-03-06
415	Verbena litoralis	Honolulu, 3830 Tantalus Dr., access road to Puu Ohia	roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum.	Preserved_Specimen	to 4-20 dm tall; corolla blue	Ray, G.	50	Oahu	BISH	1997-09-15
416	Verbena litoralis	Kewalo, back of Punchbowl	open, dry, roadside	Preserved_Specimen	1 m tall; flower - small, inconspicous, lavender	Judd, C.S.	11	Oahu	BISH	1924-11-06
417	Verbena litoralis	Lower half of Kaala Road	mesic forest	Preserved_Specimen	flowers lavender	Annable, C.R.	3727	Oahu	BISH	1997-10-15
418	Verbena litoralis	E central part of island, slope of Moaula	Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping.	Preserved_Specimen	Flowers purple.	Cuddihy, L.W.	428	Kahoolawe	BISH	1980-04-25
419	Verbena litoralis	Waikane-Schofield trail, Koolau Mts, Kahana	along trail	Preserved_Specimen	fls. lavendar	Fosberg, F.R.	10276	Oahu	BISH	1933-09-22
420	Verbena litoralis	Trail to Kaluiwaa (Sacred Falls), outside forest Reserve	At base of foot path trail.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-03-21
421	Verbena litoralis	East Maui, Haleakala Crater, Kaupo Gap	Dry volcanic gravel by trail.	Preserved_Specimen	Flws lavender-blue.	St.John, H.	21182	Maui	BISH	1945-08-27
422	Verbena litoralis	Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road	Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside..	Preserved_Specimen		Newell, C.L.	471	Hawaii	BISH	1966-08-20
423	Verbena litoralis	Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979	Pohakuloa burn 1979.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
424	Verbena litoralis	ahupuaa of Halepuaa Forest Reserve in the Puna District	Found growing on the side of the old railroad.	Preserved_Specimen		ESP Field Crew		Hawaii	BISH	1979-08-15
425	Verbena litoralis	North Kohala District, Kalala ahupua'a, Lahikiola cinder cone	In Kikuyu grassland	Preserved_Specimen		Bunch, P.M.	112	Hawaii	BISH	1989-03-13
426	Verbena litoralis	Kipapa Gulch, Wahiawa	moderately moist pineapple field	Preserved_Specimen		Hosaka, E.Y.	422	Oahu	BISH	1931-06-18
427	Verbena litoralis	Waimea Audubon Center, Waimea Valley, Oahu, Hawaii	Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions.	Preserved_Specimen	Shrub, spindly, flowers purple, on spikes.	Crago, L.M.	2005207	Oahu	BISH	2005-10-18
428	Verbena litoralis	Waiapaa Gulch, flat area near the road above	Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants.	Preserved_Specimen		Funk, E.J.	920004	Lanai	BISH	1992-01-26
429	Verbena litoralis	South Kohala: Waimea, Lalamilo	Range paddock; occasional.	Preserved_Specimen	Branching plant 1-3 ft.	Hosaka, E.Y.	2122	Hawaii	BISH	1938-06-01
430	Verbena litoralis	Kaalualu Bay near South Point	Only one individual found in waste area.	Preserved_Specimen	Herb up to one m in height with small purplish flowers.	Whistler, W.A.	42	Hawaii	BISH	1972-04-06
431	Verbena litoralis	Lawai Valley; P.T.B.G		Preserved_Specimen	Herb w/pale blue flowers. Uncommon in valley.	Herbst, D.R.	2296	Kauai	BISH	1972-01-04
432	Verbena litoralis	East Maui, Upper Kaupo Gap	in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground.	Preserved_Specimen	Flowers purple	Higashino, P.K.	885	Maui	BISH	1975-06-22
433	Verbena litoralis	Located on the Mauna Loa Strip road	about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19	Preserved_Specimen	Shrub about 4 ft. tall	Newell, C.L.	966	Hawaii	BISH	1967-06-23
434	Verbena litoralis	near Koko Head, Kaluanui Ridge,Koolau Range	guava zone; at summit	Preserved_Specimen		Egler, F.E.	3783	Oahu	BISH	1937-07-01
435	Verbena litoralis	Kaimuki, Honolulu	roadside	Preserved_Specimen		Fosberg, F.R.	8856	Oahu	BISH	1932-09-24
436	Verbena litoralis	0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline;	on rockpile, vegetation Myoporum-Chamaesyce dominant,	Preserved_Specimen		Wagner, W.L.	4688	Hawaii	BISH	1982-11-17
437	Verbena litoralis	Growing next to Kaluanui Stream	Approximately 1/2 mile from ocean.	Preserved_Specimen		Corn, C.		Oahu	BISH	1978-05-18
438	Verbena litoralis	Waianae Mts., Makaha Valley	on trail	Preserved_Specimen		Hasegawa, M.		Oahu	BISH	1932-04-03
439	Verbena litoralis	Keauhou Ranch, Kau District		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/16/2007
440	Verbena litoralis	Kalaoa, HI 96740, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	10/18/2007
441	Verbena litoralis	Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field	Roadside.	Preserved_Specimen		Wagner, W.L.	4826	Oahu	BISH	1983-04-04
442	Verbena litoralis	Honolulu, Hawaii, United States		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
443	Verbena litoralis	Growing next to Kaluanui Stream		OBSERVATION		Curtis Daehler		Oahu	HAPI	3/21/2007
444	Verbena litoralis	S half of island	Ponds	Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
445	Verbena litoralis	East Maui, floor of Haleakala Crater	1/2 mile East of Holua Cabin along trail	Preserved_Specimen		Henrickson, J.S.	3939	Maui	BISH	1969-07-20
446	Verbena litoralis	Maui		Preserved_Specimen	Flower lavender	Handy, E.S.	3320	Maui	BISH	1933-06-10
447	Verbena litoralis	Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ)	Common roadside weed.	Preserved_Specimen		Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
448	Verbena litoralis	ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision	The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica.	Preserved_Specimen		Yoshida, L.	79.053	Hawaii	BISH	1979-02-28
449	Verbena litoralis	Wahiawa		Preserved_Specimen	Erect herb, 1m; flower purple	Swarbrick, J.T.	0.62	Oahu	BISH	1976-01-07
450	Verbena litoralis	Haena		Preserved_Specimen				Kauai	BISH	1953-10-30
451	Verbena litoralis	Weed of pineapple field near northern junction of Highways 37 and 377		Preserved_Specimen	Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple.	Herbst, D.R.	795	Maui	BISH	1967-12-18
452	Verbena litoralis	Kanehoa, Waianaeuka	near top of ridge	Preserved_Specimen		Hume, E.P.	123	Oahu	BISH	1931-03-22
453	Verbena litoralis	East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store	Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus.	Preserved_Specimen		Imada, C.	200292	Maui	BISH	2002-11-19
454	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen	Herb, woody base,1.25 m high	Christophersen, E.		Oahu	BISH	1924-12-07
455	Verbena litoralis	Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility		Preserved_Specimen	Herb 1 m tall; flws light violet.	Lammers, T.G.	5313	Oahu	BISH	1983-08-27
456	Verbena litoralis	Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail		Preserved_Specimen		Wagner, W.L.	5398	Molokai	BISH	1984-07-17
457	Verbena litoralis	Maui		Preserved_Specimen		Forbes, C.N.	2036	Maui	BISH	1920-04-19
458	Verbena litoralis	Hawaii County, HI, USA		OBSERVATION		Curtis Daehler		Hawaii	HAPI	3/25/2007
459	Verbena litoralis	Pohakuloa dry area		Preserved_Specimen		Long, C.R.	4309	Hawaii	BISH	1966-12-23
460	Verbena litoralis	Maui		Preserved_Specimen		Wagner, W.L.	4783	Maui	BISH	1983-01-14
461	Verbena litoralis	Oahu		Preserved_Specimen		Rémy, J.	406	Oahu	BISH	1851-01-01
462	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	141	Lanai	BISH	1913-10-10
463	Verbena litoralis	Area within 300 m west of LZ-1. UTM coordinates 752400-2275900		Preserved_Specimen		Warren, S.	14	Kahoolawe	BISH	1987-04-09
464	Verbena litoralis	Waipio		Preserved_Specimen		Swanson, R.		Hawaii	BISH	1931-08-29
465	Verbena litoralis	Tantalus Rd		Preserved_Specimen		Forbes, A.		Oahu	BISH	1930-07-18
466	Verbena litoralis	Waikolu Valley, head of and E. rim		Preserved_Specimen		Kondo, Y.		Molokai	BISH	1962-06-18
467	Verbena litoralis	Honolulu		Preserved_Specimen		Pukui, M.K.		Oahu	BISH	1931-01-01
468	Verbena litoralis	Puu Uau, Aiea		Preserved_Specimen		Christophersen, E.		Oahu	BISH	1924-12-07
469	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
470	Verbena litoralis	Waianae Mts., South end		Preserved_Specimen		Harley, K.		Oahu	BISH	1956-05-04
471	Verbena litoralis	Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea		Preserved_Specimen		Adee, K.T.		Hawaii	BISH
472	Verbena litoralis	Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road		Preserved_Specimen		Ziegler, M.F.		Oahu	BISH	1986-11-08
473	Verbena litoralis	Kealia Salt Marsh		Preserved_Specimen		Funk, E.J.		Maui	BISH	1990-02-15
474	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
475	Verbena litoralis	Mokuleia Forest Reserve, Waianae Mts		Preserved_Specimen		Onouye, R.		Oahu	BISH	1934-02-04
476	Verbena litoralis	Valley above country club, Honolulu		Preserved_Specimen		Forbes, C.N.		Oahu	BISH	1908-07-08
477	Verbena litoralis	S half of island		Preserved_Specimen		Stokes, J.F.G.		Niihau	BISH	1912-01-01
478	Verbena litoralis	Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above		Preserved_Specimen		Sohmer, S.H.	5995	Oahu	BISH	1969-08-27
479	Verbena litoralis	Lanai		Preserved_Specimen		Munro, G.C.	292	Lanai	BISH
480	Verbena litoralis	Slopes of Mauna Kea (Wailei)		Preserved_Specimen		Forbes, C.N.	465	Hawaii	BISH	1911-08-01
481	Verbena litoralis	Kaua Trail, Waianae Mts		Preserved_Specimen		Woolford, H.A.	200	Oahu	BISH	1955-12-11
482	Verbena litoralis	Kealia trail		Preserved_Specimen		Bennett, T.	244	Oahu	BISH	1982-03-25
483	Verbena litoralis	Kualoa Pasture		Preserved_Specimen		Rock, J.F.C.	3007	Oahu	BISH	1942-01-01
484	Verbena litoralis	Kaholuamanoa, above Waimea		Preserved_Specimen		Heller, A.A.	2046	Kauai	BISH	1895-09-02
485	Verbena litoralis	Pauoa Valley		Preserved_Specimen		Garber, D.W.	339	Oahu	BISH	1920-03-21
486	Verbena litoralis	Honolulu, Aihualama		Preserved_Specimen		Russell, P.	253	Oahu	BISH	1932-07-08
487	Verbena litoralis	Crater of Haleakala		Preserved_Specimen		Forbes, C.N.	1083	Maui	BISH	1919-08-09
488	Verbena litoralis	Koele		Preserved_Specimen		Forbes, C.N.	188	Lanai	BISH	1918-06-01
489	Verbena litoralis	Maui		Preserved_Specimen		Neal, M.C.		Maui	BISH	1933-08-01
490	Verbena litoralis	unknown		Preserved_Specimen		Mann, H.	437		BISH	1864-01-01
491	Verbena litoralis	Kilauea, Kaloko Reservoir		Preserved_Specimen		Forbes, C.N.	572	Kauai	BISH	1916-10-11
492	Verbena litoralis	Palawai		Preserved_Specimen		Munro, G.C.	365	Lanai	BISH	1928-08-01
493	Verbena litoralis	Palolo Valley, middle ridge	forest	Preserved_Specimen		Christophersen, E.	1666	Oahu	BISH	1931-03-26
494	Verbena litoralis	Palaau, west side of airport	pineapple field; dry, exposed, windy.	Preserved_Specimen	Herb to sub-shrub with blue flowers	Herbst, D.R.	5092	Molokai	BISH	1974-11-11
495	Verbena litoralis	Mt. Kalena, Waianae	Lantana grove	Preserved_Specimen	herb 4 ft.	Grant, M.L.	7445	Oahu	BISH	1934-09-30
496	Verbena litoralis	Koloa, Kalaheo, upper end of Puuwai Rd.	wet roadside secondary forest; Wedelia	Preserved_Specimen	to 0.5 m herb, slender; square stem, small purple flowers	Trauernicht, C.	205	Kauai	BISH	2007-10-15
497	Verbena litoralis	Nawaimaka drainage.	Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai	Preserved_Specimen	1.5 m. tall, stems green, corolla pale pruple.	Wood, K.R.	13816	Kauai	BISH	2009-07-16
498	Verbena litoralis	ridge north of Kuapa Pond	open, hillsides	Preserved_Specimen		Gillespie, J.W.	1076	Oahu	BISH	1927-07-08
499	Verbena litoralis	Kau District: Keauhou Ranch	Along fenceline of horse corrall in lower pastures of ranch.	Preserved_Specimen	Herb, 3 ft. tall, purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
500	Verbena litoralis	stream bank below Dillingham Field wetland	grasses	Preserved_Specimen		Funk, E.J.	1105	Oahu	BISH	1990-05-08
501	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
502	Verbena litoralis	Keauhou Ranch, Kau District	Alogn fenceline of horse corrall.	Preserved_Specimen	Herb, 3 ft. tall, with purple flowers.	Davis, J.	166	Hawaii	BISH	1979-12-18
503	Verbena litoralis	Makanikahau Reservoir, Old	Dried mud	Preserved_Specimen		St.John, H.	23633	Niihau	BISH	1949-04-01
504	Verbena litoralis	Puu Huluhulu. Near the escarpment Transect (T16)	Remnant dryland community. Common subshrub among Pennisetum setaceum.	Preserved_Specimen	Common subshrub.	Takeuchi, W.N.	5714	Hawaii	BISH	1989-12-26
505	Verbena litoralis	Kaluanui Flats	Pineapple fields	Preserved_Specimen	Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas.	Spence, G.	369	Lanai	BISH	1973-10-24
506	Verbena litoralis	Mauna Kea (Puu Laau Road)	In open Mamane-Naio forest.	Preserved_Specimen		Engilis Jr., A.	39	Hawaii	BISH	1987-08-19
507	Verbena litoralis	Keanakolu, Hilo Forest Reserve	Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus.	Preserved_Specimen		Higashino, P.K.	6965	Hawaii	BISH	1978-03-01
508	Verbena litoralis	Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost	in open sunny area.	Preserved_Specimen		Herat, T.	559	Hawaii	BISH	1974-06-12
509	Verbena litoralis	North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary	Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa	Preserved_Specimen		Close, B.	109426	Hawaii	BISH	1994-10-29
510	Verbena litoralis	Peacock Flat, Waianae Mts	roadside, dry forest	Preserved_Specimen	herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue.	Spence, G.	209	Oahu	BISH	1973-09-06
511	Verbena litoralis	Puu Laau	Weed growing near the hunter's cabin.	Preserved_Specimen		Herbst, D.R.	5210	Hawaii	BISH	1975-01-18
512	Verbena litoralis	Kahaluu; Kailua; Kapua	Koa forest.	Preserved_Specimen		Adee, K.T.		Hawaii	BISH
513	Verbena litoralis	Puuwaawaa; track to 1986 burn area (from Ranch HQ)	4WD access track to burn area.	Preserved_Specimen	Common roadside weed.	Takeuchi, W.N.	5752	Hawaii	BISH	1989-05-01
514	Verbena litoralis	Kawa side of Kahana Valley, pass the main gate and the marshy area	along jeep trail, open areas	Preserved_Specimen	common weed, ~4', with light blue flowers	Herat, T.	158	Oahu	BISH	1972-08-14
515	Verbena litoralis	1750' Field 123, Honokaa GSugar Co. Hamakua District	Common weed at field edge, and on jeep roads through field, not present inside mature stand.	Preserved_Specimen	Perennial root, plants 30- 48 tall, corolla pale blue	Tomich, P.Q.	911	Hawaii	BISH	1960-03-18
516	Verbena litoralis	Kahuku, Ocean View Estates,	open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive.	Preserved_Specimen	Common roadside weed with blue flowers.	Herbst, D.R.	5068	Hawaii	BISH	1974-10-09
517	Verbena litoralis	Kilauea	weed in abandoned sorghum field	Preserved_Specimen		Herbst, D.R.	5244	Kauai	BISH	1975-02-27
518	Verbena litoralis	Laie	clay, pasture	Preserved_Specimen	herb 1-6', tall slender stems	Guinto, J.	27	Oahu	BISH	1986-03-06
519	Verbena litoralis	Honolulu, 3830 Tantalus Dr., access road to Puu Ohia	roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum.	Preserved_Specimen	to 4-20 dm tall; corolla blue	Ray, G.	50	Oahu	BISH	1997-09-15
520	Verbena litoralis	Kewalo, back of Punchbowl	open, dry, roadside	Preserved_Specimen	1 m tall; flower - small, inconspicous, lavender	Judd, C.S.	11	Oahu	BISH	1924-11-06

Verbena litoralis

Kunth (1818)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Verbena

‘ōwī [owi], ha‘uoi (Ni‘ihau via St. John), oī [oi], ōwī, vervain

Description

Historical Distribution

Images

Synonyms (32)

Uses and Culture

USES

Natural History

Statewide Status

Island Status

Dispersal Agents

Pollinators

Specimens

Conservation Status

Notes

Bibliography

Occurrences

Verbena litoralis

Kunth (1818)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Verbena

‘ōwī [owi], ha‘uoi (Ni‘ihau via St. John), oī [oi], ōwī, vervain

Description

Historical Distribution

Images

Synonyms (32)

Uses and Culture

USES

Natural History

Statewide Status

Island Status

Dispersal Agents

Pollinators

Specimens

Conservation Status

Notes

Bibliography

Occurrences

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