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Verbena litoralis

Kunth (1818)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Verbena

‘ōwī [owi], ha‘uoi (Ni‘ihau via St. John), oī [oi], ōwī, vervain

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Description

Key Characters:

Growth Form: Erect or ascending perennial herbs 4–20 dm tall, sometimes somewhat woody toward base.

Stems: Stems few to many-branched, glabrous or sparsely strigillose.

Roots:

Leaves: Leaves simple. Opposite and decussate (pairs of leaves alternating 90 degrees between nodes). Blades lanceolate to oblanceolate, 3–10 cm long, 1–1.5 cm wide. Apex acute. Base attenuate. Surfaces finely scaberulous. Margins coarsely serrate. Veins prominent. Sessile. Stipules absent.

Flowers: Flowers numerous in several slender, terminal spikes 3–12 cm long, sparsely short–pilose throughout; Bracts subulate, shorter than to as long as the calyx; short–pilose. Flowers bisexual (perfect), alternate, nearly zygomorphic. Calyx sometimes purple, 1.5–2.3 mm long. Corolla usually blue, occasionally pale blue, lavender, or purple, ca. 2–3 mm long, 5-lobed, imbricate. Stamens 4, in 2 pairs, inserted on and included in corolla tube; anthers dithecal, opening by longitudinal slits. Ovary superior, 4-lobed, 2-carpellate, 4-celled, placentation axile; ovules 1 per cell, attached laterally or near base, erect, anatropous; style 2-lobed, the lobes dissimilar, only 1 stigmatic.

Fruit: Schizocarps readily separating at maturity into 4 crustaceous nutlets; pericarp hard and dry. Seeds 1 per nutlet; oblong; triangular in cross section; endosperm essentially absent.

Ploidy: 2n = 28; 56

Habitat: Occurring in dry to wet; disturbed habitats.

Elevation Range: 10–2,280 m.

Historical Distribution

Images

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Synonyms (32)

Verbena affinis M.Martens & Galeotti (1844)
Verbena bonariensis (Misapplied)
Verbena bonariensis var. litoralis (Kunth) Hook. ex Müll.Berol. (1845)
Verbena brasiliensis var. subglabrata Moldenke (1950)
Verbena cajamarcensis Binder ex Molinari (2015)
Verbena caracasana Kunth (1818)
Verbena carolina var. glabra Moldenke (1959)
Verbena gentryi Moldenke (1941)
Verbena glabrata var. tenuispicata Moldenke (1967)
Verbena integrifolia Sessé & Moc. (1886)
Verbena integrifolia f. albiflora Moldenke (1968)
Verbena lanceolata Willd. ex Spreng. (1825)
Verbena litoralis f. albiflora (Moldenke) Moldenke (1979)
Verbena litoralis f. magnifolia Moldenke (1977)
Verbena litoralis var. albiflora Moldenke (1940)
Verbena litoralis var. caracasana (Kunth) Briq. (1904)
Verbena litoralis var. glabrior Benth. (1846)
Verbena litoralis var. leptostachya Schauer (1847)
Verbena litoralis var. litoralis
Verbena litoralis var. melanopotamica Hauman (1913)
Verbena litoralis var. portoricensis Moldenke (1982)
Verbena litoralis var. pycnostachya Schauer (1847)
Verbena litoralis var. subglabrata (Moldenke) N.O'Leary & Múlgura (2012)
Verbena longifolia M.Martens & Galeotti (1844)
Verbena longifolia f. albiflora Moldenke (1961)
Verbena longifolia var. pubescens Moldenke (1966)
Verbena minutiflora var. peruviana Moldenke (1981)
Verbena nudiflora Nutt. ex Turcz. (1863)
Verbena paucifolia Turcz. (1863)
Verbena sedula Moldenke (1955)
Verbena sedula var. darwinii Moldenke (1968)
Verbena sedula var. fournieri Moldenke (1969)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Ni'ihau(Incl. Lehua) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Kaho'olawe Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Notes

  • Description digitized by Ikaika Mendez
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • Has been used medicinally in Hawai'i especially as a mash applied to cuts and bruises and also to sprained and fractured areas.
  • in 1837, specific locality unknown

Bibliography

Name Published In: F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 2: 276 (1818)

Other References

Wagner et al. 1990:1325 (Mi, Ni, K, O, Mo, L, M, Ka, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
2 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
3 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
4 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
5 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
6 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
7 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
8 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
9 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
10 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
11 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
12 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
13 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
14 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
15 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
16 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
17 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
18 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
19 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
20 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
21 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
22 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
23 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
24 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
25 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
26 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
27 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
28 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
29 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
30 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
31 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
32 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
33 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
34 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
35 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
36 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
37 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
38 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
39 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
40 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
41 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
42 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
43 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
44 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
45 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
46 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
47 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
48 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
49 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
50 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
51 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
52 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
53 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
54 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
55 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
56 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
57 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
58 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
59 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
60 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
61 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
62 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
63 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
64 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
65 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
66 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
67 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
68 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
69 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
70 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
71 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
72 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
73 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
74 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
75 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
76 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
77 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
78 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
79 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
80 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
81 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
82 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
83 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
84 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
85 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
86 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
87 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
88 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
89 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
90 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
91 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
92 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
93 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
94 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
95 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
96 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
97 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
98 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
99 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
100 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
101 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
102 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
103 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
104 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
105 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
106 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
107 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
108 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
109 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
110 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
111 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
112 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
113 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
114 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
115 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
116 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
117 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
118 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
119 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
120 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
121 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
122 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
123 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
124 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
125 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
126 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
127 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
128 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
129 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
130 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
131 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
132 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
133 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
134 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
135 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
136 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
137 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
138 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
139 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
140 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
141 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
142 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
143 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
144 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
145 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
146 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
147 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
148 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
149 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
150 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
151 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
152 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
153 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
154 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
155 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
156 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
157 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
158 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
159 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
160 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
161 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
162 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
163 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
164 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
165 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
166 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
167 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
168 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
169 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
170 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
171 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
172 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
173 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
174 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
175 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
176 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
177 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
178 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
179 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
180 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
181 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
182 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
183 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
184 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
185 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
186 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
187 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
188 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
189 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
190 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
191 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
192 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
193 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
194 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
195 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
196 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
197 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
198 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
199 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
200 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
201 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
202 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
203 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
204 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
205 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
206 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
207 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
208 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
209 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
210 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
211 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
212 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
213 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
214 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
215 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
216 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
217 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
218 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
219 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
220 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
221 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
222 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
223 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
224 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
225 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
226 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
227 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
228 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
229 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
230 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
231 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
232 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
233 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
234 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
235 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
236 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
237 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
238 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
239 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
240 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
241 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
242 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
243 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
244 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
245 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
246 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
247 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
248 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
249 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
250 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
251 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
252 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
253 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
254 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
255 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
256 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
257 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
258 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
259 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
260 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
261 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
262 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
263 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
264 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
265 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
266 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
267 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
268 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
269 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
270 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
271 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
272 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
273 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
274 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
275 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
276 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
277 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
278 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
279 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
280 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
281 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
282 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
283 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
284 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
285 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
286 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
287 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
288 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
289 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
290 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
291 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
292 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
293 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
294 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
295 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
296 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
297 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
298 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
299 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
300 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
301 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
302 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
303 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
304 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
305 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
306 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
307 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
308 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
309 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
310 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
311 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
312 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
313 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
314 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
315 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
316 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
317 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
318 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
319 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
320 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
321 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
322 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
323 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
324 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
325 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
326 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
327 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
328 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
329 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
330 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
331 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
332 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
333 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
334 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
335 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
336 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
337 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
338 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
339 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
340 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
341 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
342 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
343 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
344 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
345 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
346 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
347 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
348 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
349 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
350 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
351 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
352 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
353 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
354 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
355 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
356 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
357 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
358 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
359 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
360 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
361 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
362 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
363 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
364 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
365 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
366 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
367 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
368 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
369 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
370 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
371 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
372 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
373 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
374 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
375 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
376 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
377 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
378 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
379 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
380 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
381 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
382 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
383 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
384 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
385 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
386 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
387 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
388 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
389 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
390 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
391 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
392 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
393 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
394 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
395 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
396 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
397 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
398 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
399 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
400 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
401 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
402 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
403 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
404 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
405 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
406 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
407 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
408 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
409 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
410 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
411 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
412 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
413 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
414 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
415 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
416 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
417 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
418 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
419 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
420 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
421 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
422 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
423 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
424 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
425 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
426 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
427 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
428 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
429 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
430 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
431 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
432 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
433 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
434 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
435 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
436 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
437 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
438 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
439 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
440 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
441 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
442 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
443 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
444 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
445 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
446 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
447 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
448 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
449 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
450 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
451 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
452 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
453 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
454 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
455 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
456 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
457 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
458 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
459 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
460 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
461 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
462 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
463 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
464 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
465 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
466 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
467 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
468 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
469 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
470 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
471 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
472 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
473 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
474 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
475 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
476 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
477 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
478 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
479 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
480 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
481 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
482 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
483 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
484 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
485 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
486 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
487 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
488 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
489 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
490 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
491 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
492 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
493 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
494 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
495 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
496 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
497 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
498 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
499 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
500 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
501 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
502 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
503 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
504 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
505 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
506 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
507 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
508 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
509 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
510 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
511 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
512 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
513 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
514 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
515 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
516 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
517 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
518 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
519 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
520 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
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