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Verbena litoralis

Kunth (1818)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Verbena

‘ōwī [owi], ha‘uoi (Ni‘ihau via St. John), oī [oi], ōwī, vervain

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Description

Key Characters:

Growth Form: Erect or ascending perennial herbs 4–20 dm tall, sometimes somewhat woody toward base.

Stems: Stems few to many-branched, glabrous or sparsely strigillose.

Roots:

Leaves: Leaves simple. Opposite and decussate (pairs of leaves alternating 90 degrees between nodes). Blades lanceolate to oblanceolate, 3–10 cm long, 1–1.5 cm wide. Apex acute. Base attenuate. Surfaces finely scaberulous. Margins coarsely serrate. Veins prominent. Sessile. Stipules absent.

Flowers: Flowers numerous in several slender, terminal spikes 3–12 cm long, sparsely short–pilose throughout; Bracts subulate, shorter than to as long as the calyx; short–pilose. Flowers bisexual (perfect), alternate, nearly zygomorphic. Calyx sometimes purple, 1.5–2.3 mm long. Corolla usually blue, occasionally pale blue, lavender, or purple, ca. 2–3 mm long, 5-lobed, imbricate. Stamens 4, in 2 pairs, inserted on and included in corolla tube; anthers dithecal, opening by longitudinal slits. Ovary superior, 4-lobed, 2-carpellate, 4-celled, placentation axile; ovules 1 per cell, attached laterally or near base, erect, anatropous; style 2-lobed, the lobes dissimilar, only 1 stigmatic.

Fruit: Schizocarps readily separating at maturity into 4 crustaceous nutlets; pericarp hard and dry. Seeds 1 per nutlet; oblong; triangular in cross section; endosperm essentially absent.

Ploidy: 2n = 28; 56

Habitat: Occurring in dry to wet; disturbed habitats.

Elevation Range: 10–2,280 m.

Historical Distribution

Images

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Synonyms (32)

Verbena affinis M.Martens & Galeotti (1844)
Verbena bonariensis (Misapplied)
Verbena bonariensis var. litoralis (Kunth) Hook. ex Müll.Berol. (1845)
Verbena brasiliensis var. subglabrata Moldenke (1950)
Verbena cajamarcensis Binder ex Molinari (2015)
Verbena caracasana Kunth (1818)
Verbena carolina var. glabra Moldenke (1959)
Verbena gentryi Moldenke (1941)
Verbena glabrata var. tenuispicata Moldenke (1967)
Verbena integrifolia Sessé & Moc. (1886)
Verbena integrifolia f. albiflora Moldenke (1968)
Verbena lanceolata Willd. ex Spreng. (1825)
Verbena litoralis f. albiflora (Moldenke) Moldenke (1979)
Verbena litoralis f. magnifolia Moldenke (1977)
Verbena litoralis var. albiflora Moldenke (1940)
Verbena litoralis var. caracasana (Kunth) Briq. (1904)
Verbena litoralis var. glabrior Benth. (1846)
Verbena litoralis var. leptostachya Schauer (1847)
Verbena litoralis var. litoralis
Verbena litoralis var. melanopotamica Hauman (1913)
Verbena litoralis var. portoricensis Moldenke (1982)
Verbena litoralis var. pycnostachya Schauer (1847)
Verbena litoralis var. subglabrata (Moldenke) N.O'Leary & Múlgura (2012)
Verbena longifolia M.Martens & Galeotti (1844)
Verbena longifolia f. albiflora Moldenke (1961)
Verbena longifolia var. pubescens Moldenke (1966)
Verbena minutiflora var. peruviana Moldenke (1981)
Verbena nudiflora Nutt. ex Turcz. (1863)
Verbena paucifolia Turcz. (1863)
Verbena sedula Moldenke (1955)
Verbena sedula var. darwinii Moldenke (1968)
Verbena sedula var. fournieri Moldenke (1969)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Ni'ihau(Incl. Lehua) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Kaho'olawe Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Notes

  • Description digitized by Ikaika Mendez
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • Has been used medicinally in Hawai'i especially as a mash applied to cuts and bruises and also to sprained and fractured areas.
  • in 1837, specific locality unknown

Bibliography

Name Published In: F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 2: 276 (1818)

Other References

Wagner et al. 1990:1325 (Mi, Ni, K, O, Mo, L, M, Ka, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
2 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
3 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
4 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
5 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
6 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
7 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
8 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
9 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
10 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
11 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
12 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
13 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
14 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
15 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
16 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
17 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
18 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
19 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
20 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
21 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
22 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
23 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
24 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
25 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
26 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
27 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
28 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
29 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
30 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
31 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
32 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
33 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
34 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
35 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
36 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
37 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
38 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
39 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
40 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
41 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
42 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
43 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
44 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
45 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
46 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
47 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
48 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
49 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
50 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
51 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
52 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
53 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
54 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
55 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
56 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
57 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
58 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
59 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
60 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
61 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
62 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
63 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
64 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
65 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
66 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
67 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
68 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
69 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
70 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
71 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
72 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
73 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
74 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
75 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
76 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
77 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
78 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
79 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
80 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
81 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
82 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
83 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
84 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
85 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
86 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
87 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
88 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
89 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
90 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
91 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
92 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
93 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
94 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
95 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
96 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
97 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
98 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
99 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
100 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
101 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
102 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
103 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
104 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
105 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
106 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
107 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
108 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
109 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
110 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
111 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
112 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
113 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
114 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
115 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
116 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
117 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
118 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
119 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
120 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
121 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
122 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
123 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
124 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
125 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
126 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
127 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
128 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
129 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
130 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
131 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
132 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
133 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
134 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
135 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
136 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
137 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
138 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
139 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
140 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
141 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
142 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
143 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
144 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
145 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
146 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
147 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
148 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
149 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
150 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
151 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
152 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
153 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
154 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
155 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
156 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
157 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
158 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
159 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
160 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
161 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
162 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
163 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
164 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
165 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
166 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
167 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
168 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
169 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
170 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
171 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
172 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
173 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
174 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
175 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
176 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
177 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
178 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
179 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
180 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
181 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
182 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
183 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
184 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
185 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
186 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
187 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
188 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
189 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
190 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
191 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
192 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
193 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
194 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
195 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
196 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
197 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
198 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
199 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
200 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
201 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
202 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
203 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
204 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
205 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
206 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
207 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
208 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
209 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
210 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
211 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
212 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
213 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
214 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
215 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
216 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
217 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
218 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
219 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
220 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
221 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
222 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
223 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
224 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
225 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
226 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
227 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
228 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
229 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
230 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
231 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
232 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
233 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
234 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
235 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
236 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
237 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
238 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
239 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
240 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
241 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
242 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
243 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
244 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
245 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
246 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
247 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
248 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
249 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
250 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
251 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
252 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
253 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
254 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
255 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
256 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
257 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
258 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
259 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
260 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
261 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
262 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
263 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
264 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
265 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
266 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
267 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
268 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
269 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
270 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
271 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
272 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
273 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
274 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
275 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
276 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
277 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
278 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
279 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
280 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
281 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
282 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
283 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
284 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
285 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
286 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
287 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
288 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
289 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
290 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
291 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
292 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
293 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
294 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
295 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
296 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
297 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
298 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
299 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
300 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
301 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
302 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
303 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
304 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
305 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
306 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
307 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
308 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
309 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
310 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
311 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
312 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
313 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
314 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
315 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
316 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
317 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
318 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
319 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
320 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
321 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
322 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
323 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
324 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
325 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
326 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
327 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
328 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
329 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
330 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
331 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
332 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
333 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
334 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
335 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
336 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
337 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
338 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
339 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
340 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
341 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
342 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
343 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
344 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
345 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
346 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
347 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
348 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
349 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
350 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
351 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
352 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
353 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
354 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
355 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
356 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
357 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
358 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
359 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
360 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
361 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
362 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
363 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
364 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
365 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
366 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
367 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
368 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
369 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
370 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
371 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
372 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
373 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
374 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
375 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
376 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
377 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
378 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
379 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
380 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
381 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
382 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
383 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
384 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
385 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
386 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
387 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
388 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
389 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
390 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
391 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
392 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
393 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
394 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
395 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
396 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
397 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
398 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
399 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
400 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
401 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
402 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
403 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
404 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
405 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
406 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
407 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
408 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
409 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
410 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
411 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
412 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
413 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
414 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
415 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
416 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
417 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
418 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
419 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
420 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
421 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
422 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
423 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
424 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
425 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
426 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
427 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
428 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
429 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
430 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
431 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
432 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
433 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
434 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
435 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
436 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
437 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
438 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
439 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
440 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
441 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
442 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
443 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
444 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
445 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
446 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
447 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
448 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
449 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
450 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
451 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
452 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
453 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
454 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
455 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
456 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
457 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
458 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
459 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
460 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
461 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
462 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
463 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
464 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
465 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
466 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
467 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
468 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
469 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
470 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
471 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
472 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
473 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
474 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
475 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
476 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
477 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
478 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
479 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
480 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
481 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
482 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
483 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
484 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
485 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
486 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
487 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
488 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
489 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
490 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
491 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
492 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
493 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
494 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
495 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
496 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
497 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
498 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
499 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
500 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
501 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
502 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
503 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
504 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
505 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
506 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
507 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
508 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
509 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
510 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
511 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
512 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
513 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
514 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
515 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
516 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
517 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
518 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
519 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
520 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
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