Verbena litoralis

Kunth (1818)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Verbena

‘ōwī [owi], ha‘uoi (Ni‘ihau via St. John), oī [oi], ōwī, vervain

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Description

Key Characters:

Growth Form: Erect or ascending perennial herbs 4–20 dm tall, sometimes somewhat woody toward base.

Stems: Stems few to many-branched, glabrous or sparsely strigillose.

Roots:

Leaves: Leaves simple. Opposite and decussate (pairs of leaves alternating 90 degrees between nodes). Blades lanceolate to oblanceolate, 3–10 cm long, 1–1.5 cm wide. Apex acute. Base attenuate. Surfaces finely scaberulous. Margins coarsely serrate. Veins prominent. Sessile. Stipules absent.

Flowers: Flowers numerous in several slender, terminal spikes 3–12 cm long, sparsely short–pilose throughout; Bracts subulate, shorter than to as long as the calyx; short–pilose. Flowers bisexual (perfect), alternate, nearly zygomorphic. Calyx sometimes purple, 1.5–2.3 mm long. Corolla usually blue, occasionally pale blue, lavender, or purple, ca. 2–3 mm long, 5-lobed, imbricate. Stamens 4, in 2 pairs, inserted on and included in corolla tube; anthers dithecal, opening by longitudinal slits. Ovary superior, 4-lobed, 2-carpellate, 4-celled, placentation axile; ovules 1 per cell, attached laterally or near base, erect, anatropous; style 2-lobed, the lobes dissimilar, only 1 stigmatic.

Fruit: Schizocarps readily separating at maturity into 4 crustaceous nutlets; pericarp hard and dry. Seeds 1 per nutlet; oblong; triangular in cross section; endosperm essentially absent.

Ploidy: 2n = 28; 56

Habitat: Occurring in dry to wet; disturbed habitats.

Elevation Range: 10–2,280 m.

Historical Distribution

Images

Synonyms (32)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Ni'ihau(Incl. Lehua) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Kaho'olawe Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Notes

  • Description digitized by Ikaika Mendez
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • Has been used medicinally in Hawai'i especially as a mash applied to cuts and bruises and also to sprained and fractured areas.
  • in 1837, specific locality unknown

Bibliography

Name Published In: F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 2: 276 (1818)

Other References

Wagner et al. 1990:1325 (Mi, Ni, K, O, Mo, L, M, Ka, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
2 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
3 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
4 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
5 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
6 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
7 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
8 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
9 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
10 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
11 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
12 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
13 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
14 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
15 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
16 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
17 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
18 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
19 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
20 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
21 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
22 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
23 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
24 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
25 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
26 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
27 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
28 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
29 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
30 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
31 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
32 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
33 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
34 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
35 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
36 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
37 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
38 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
39 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
40 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
41 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
42 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
43 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
44 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
45 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
46 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
47 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
48 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
49 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
50 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
51 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
52 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
53 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
54 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
55 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
56 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
57 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
58 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
59 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
60 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
61 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
62 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
63 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
64 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
65 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
66 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
67 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
68 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
69 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
70 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
71 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
72 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
73 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
74 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
75 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
76 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
77 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
78 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
79 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
80 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
81 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
82 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
83 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
84 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
85 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
86 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
87 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
88 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
89 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
90 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
91 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
92 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
93 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
94 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
95 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
96 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
97 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
98 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
99 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
100 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
101 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
102 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
103 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
104 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
105 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
106 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
107 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
108 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
109 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
110 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
111 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
112 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
113 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
114 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
115 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
116 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
117 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
118 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
119 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
120 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
121 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
122 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
123 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
124 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
125 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
126 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
127 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
128 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
129 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
130 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
131 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
132 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
133 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
134 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
135 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
136 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
137 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
138 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
139 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
140 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
141 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
142 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
143 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
144 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
145 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
146 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
147 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
148 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
149 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
150 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
151 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
152 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
153 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
154 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
155 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
156 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
157 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
158 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
159 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
160 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
161 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
162 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
163 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
164 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
165 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
166 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
167 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
168 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
169 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
170 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
171 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
172 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
173 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
174 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
175 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
176 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
177 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
178 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
179 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
180 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
181 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
182 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
183 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
184 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
185 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
186 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
187 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
188 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
189 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
190 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
191 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
192 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
193 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
194 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
195 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
196 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
197 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
198 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
199 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
200 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
201 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
202 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
203 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
204 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
205 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
206 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
207 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
208 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
209 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
210 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
211 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
212 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
213 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
214 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
215 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
216 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
217 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
218 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
219 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
220 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
221 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
222 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
223 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
224 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
225 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
226 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
227 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
228 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
229 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
230 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
231 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
232 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
233 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
234 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
235 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
236 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
237 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
238 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
239 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
240 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
241 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
242 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
243 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
244 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
245 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
246 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
247 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
248 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
249 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
250 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
251 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
252 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
253 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
254 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
255 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
256 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
257 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
258 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
259 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
260 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
261 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
262 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
263 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
264 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
265 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
266 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
267 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
268 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
269 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
270 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
271 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
272 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
273 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
274 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
275 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
276 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
277 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
278 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
279 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
280 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
281 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
282 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
283 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
284 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
285 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
286 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
287 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
288 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
289 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
290 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
291 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
292 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
293 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
294 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
295 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
296 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
297 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
298 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
299 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
300 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
301 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
302 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
303 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
304 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
305 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
306 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
307 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
308 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
309 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
310 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
311 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
312 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
313 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
314 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
315 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
316 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
317 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
318 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
319 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
320 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
321 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
322 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
323 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
324 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
325 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
326 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
327 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
328 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
329 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
330 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
331 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
332 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
333 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
334 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
335 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
336 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
337 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
338 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
339 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
340 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
341 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
342 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
343 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
344 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
345 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
346 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
347 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
348 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
349 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
350 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
351 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
352 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
353 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
354 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
355 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
356 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
357 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
358 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
359 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
360 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
361 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
362 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
363 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
364 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
365 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
366 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
367 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
368 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
369 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
370 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
371 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
372 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
373 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
374 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
375 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
376 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
377 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
378 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
379 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
380 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
381 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
382 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
383 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
384 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
385 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
386 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
387 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
388 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
389 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
390 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
391 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
392 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
393 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
394 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
395 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
396 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
397 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
398 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
399 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
400 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
401 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
402 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
403 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
404 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
405 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
406 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
407 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
408 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
409 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
410 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
411 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
412 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
413 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
414 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
415 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
416 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
417 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
418 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
419 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
420 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
421 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
422 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
423 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
424 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
425 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
426 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
427 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
428 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
429 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
430 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
431 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
432 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
433 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
434 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
435 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
436 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
437 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
438 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
439 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
440 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
441 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
442 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
443 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
444 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
445 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
446 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
447 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
448 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
449 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
450 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
451 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
452 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
453 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
454 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
455 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
456 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
457 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
458 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
459 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
460 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
461 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
462 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
463 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
464 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
465 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
466 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
467 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
468 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
469 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
470 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
471 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
472 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
473 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
474 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
475 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
476 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
477 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
478 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
479 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
480 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
481 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
482 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
483 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
484 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
485 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
486 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
487 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
488 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
489 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
490 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
491 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
492 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
493 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
494 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
495 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
496 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
497 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
498 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
499 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
500 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
501 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
502 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
503 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
504 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
505 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
506 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
507 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
508 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
509 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
510 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
511 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
512 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
513 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
514 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
515 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
516 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
517 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
518 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
519 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
520 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15